连珠蕨属 liang zhu jue shu
Zhang Xianchun (张宪春)1;
Pseudodrynaria (C. Christensen) C. Christensen.
Plants epiphytic, epilithic, or terrestrial. Rhizome thick, short to long creeping; vascular bundles 20100, arranged in 1 or 2 flattened circles in cross section with conspicuous dorsal invaginations or protrusions; sclerenchyma strands absent. Rhizome scales appressed or spreading, pseudopeltate or rarely peltate, margin toothed or ciliate with 1- or 2-celled glandular projections. Fronds not articulate, monomorphic, usually internally dimorphic, sessile with a dilated base, frond bases imbricate or separate, forming individual nests, rachises not persistent; lamina deeply pinnatifid or subpinnate, with conspicuous nectaries situated below junctions of rachis and costae, or of costae and veins. Pinnae abscissile[??spelling] from costa and from each other by a line of abscission between costa and base of sinus, gradually smaller toward frond apex, entire, apical pinna present. Venation highly complex, with main areoles delimited by veins and connecting veins, filled with many small areoles containing excurrent and recurrent free veinlets, each veinlet terminating in a hydathode. Fertile parts similar to sterile or usually narrower. Sori small, in rows along connecting veins or veinlets, or distinctly enlarged to soral[??spelling] patches, in one row between midrib and margin. Sporangia glabrous or sometimes with 13 acicular hairs. Spores with spines or small globules. [??x] n = 36, 37.
About 31 species restricted to tropical Asia, from Himalaya to Taiwan, most abundantly in Malesia; two species in China.
Plants of Aglaomorpha mainly occur in tropical forests, forming large nests around tree trunks or on rocks. Aglaomorpha is like Drynaria, but the fronds are internally dimorphic. Photinopteris, possessing several unique characters, is not included here; otherwise, the delimitation is according to Roos (1985 [??citation]).
1a...... Fertile pinnae similar to sterile pinnae ................................................................................................. 1. A. coronans
1b..... Fertile pinnae distinctly contracted .................................................................................................. 2. A. meyeniana
崖姜 ya jiang
Polypodium coronans Wallich ex Mettenius, Abh. Senckenberg. Naturf. Ges. (Polypodium): 121. 1857; Drynaria conjugata Beddome (1870), not T. Moore (1862); D. coronans (Wallich ex Mettenius) J. Smith ex T. Moore; D. esquirolii C. Christensen; Pleopeltis coronans (Wallich ex Mettenius) Alderwerelt; Pseudodrynaria coronans (Wallich ex Mettenius) Ching; Polypodium conjugatum Baker (1868), not Poiret (1804), nor Kaulfuss (1827).
Rhizome 23 cm in diam. or more; dilated from imbricate [frond] bases; rachises not persistent. Rhizome scales spreading, pseudopeltate, 520 Χ 0.51 mm, basal auricles short, margin toothed, apex narrowly acuminate to shortly filiform; midrib absent. Fronds monomorphic or with fertile parts slightly narrowed, sessile, dilated base lobed, upward pinnatifid up to 0.2 cm from rachis, 70170 Χ 2045 cm, glabrous; apical pinna present. Pinnae without basal constriction, 1535 Χ 1.55 cm, margin entire, apex acute to acuminate. Sori usually present on all pinnae, in several rows between midrib and margin, one in each main areole, elongated, 13 mm in largest diameter, slightly sunken. Sporangia glabrous. Spores with spines and verrucate[??verrucose].
Epiphytic, forming a ring-shaped basket around tree trunks, or epilithic, primary and secondary forests; 1001900 m. Fujian, Guangdong, Guangxi, Hainan, Guizhou, Taiwan, S Xizang, S Yunnan [Indochina [??countries], Myanmar, India, Nepal, Peninsular Malaysia, Ryukyu Islands].
The name Phymatodes coronans Wallich ex C. Presl (Tent. Pterid.: 198. 1836) is an invalid nom. nud.
连珠蕨 liang zhu jue
Dryostachyum meyenianum (Schott) Brause [??spelling]; Pleopeltis meyeniana (Schott) Alderwerelt; Polypodium meyenianum (Schott) Hooker; Psygmium elegans C. Presl.
Rhizome shortly creeping, 23 cm in diam. or more; dilated frond bases imbricate; rachises not persistent. Rhizome scales spreading, pseudopeltate, 615 Χ 0.41(1.3) mm, basal auricles short, apex narrowly acuminate to long-filiform, strongly dentate; midrib absent. Fronds internally dimorphic, sessile, dilated base lobed, upward pinnatifid, 3590 Χ 1530 cm, glabrous or abscission vein with tufts of very short acicular hairs, apical pinna present. Sterile pinnae without basal constriction, 7.515 Χ 1.53.5 cm, apex rounded or acute to acuminate. Fully fertile pinnae in upper 2/3 of frond, distinctly narrowed, up to 520 Χ 0.40.8 cm, contracted between sori. Sorial[??sp] patches in one row along pinna midrib, protruding and beadlike, round, 13 mm wide. Sporangia glabrous. Spores verrucate [??verrrucose].
Epiphytic, forming a ring-shaped basket around tree trunks, or epilithic, or terrestrial, usually in exposed areas in primary forests; 4001600 m. Taiwan [Philippines].
顶育蕨属 ding yu jue shu
Zhang Xianchun (张宪春)1;
Like Drynaria but differs from it in: fronds stipitate, without basal fronds or expanded lamina at lower part, fertile pinnae very narrow, elongate, sori acrostichous. [??description not parallel with others]
One species: China, Indochina, Indonesia, Malaysia, Philippines, Thailand.
Several monotypic genera were segregated from Photinopteris principally by the difference in the form of the fertile pinnae. These were all included within Aglaomorpha by the monographer Roos (Phylog. Syst. Drynar. Polyp. [Utrecht] 227. 1985). Photinopteris is maintained here pending further research, especially molecular evidence.
顶育蕨 ding yu jue
Acrostichum acuminatum Willdenow, Sp. Pl., ed. 4, 5: 116. 1810; Aglaomorpha acuminata (Willdenow) Hovenkamp; A. speciosa (Blume) M. C. Roos; Lomaria speciosa Blume; Photinopteris speciosa (Blume) C. Presl; Polypodium speciosum (Blume) H. Christ (1897), not Blume (1828), nor Meyen (1834).
Rhizome glaucous, long-creeping, 720 mm in diam.; scales brown, linear-lanceolate, 310 Χ 0.21.2 mm, base auriculate, margin dentate to ciliate, apex narrowly acuminate to filiform-subulate. Fronds internally dimorphic, stalked; stipe 1035 cm, with two rows of rudimentary pinnae; lamina pinnate, 30()4590 Χ 1330 cm, leathery, glabrous or with sparse hairs up to 3 mm, apical pinna present, nectaries on small extensions of pinna base at basiscopic side, sometimes also at acroscopic side; sterile pinnae ovate, 730 Χ 3.510 cm, base cuneate, apex acuminate to caudate, stalks up to 1 cm; fertile pinnae in upper 2/3 of frond, few to several pairs, strongly narrowed, linear, 1027 cm Χ 47 mm. Sori (coenosori) linear, covering whole abaxial surface of fertile pinnae except for costa and near margin. Spores with small globules.
Epilithic, climbing on limestone cliffs in exposed or shaded places in tropical rain forests, forming large populations, but locally uncommon; 13001400 m. S Yunnan [Indochina [??countries], Indonesia, Malaysia, Philippines, Thailand].
节肢蕨属 jie zhi jue shu
Pleopeltis sect. Arthromeris T. Moore, Index Filicum 78. 1857.
Plants, epiphytic or terrestrial, medium-sized. Rhizome long-creeping, densely scaly or covered with whitish bloom and sparse scales; scales whitish, brown, or black, pseudopeltate, lanceolate-attenuate, margin entire to shallowly toothed or ciliate. Fronds remote, monomorphic, articulate to rhizome on short scaly phyllopodes. Stipe straw-colored or brown, glabrous or pubescent. Lamina imparipinnate or simple. Lateral pinnae opposite or subopposite, rarely alternate, articulate to rachis, mostly sessile or subsessile, usually oblong-lanceolate, margin entire and cartilaginous, apex acuminate; costae and lateral veins distinct; veinlets obscure, reticulate to form irregularly shaped areoles with simple or forked included veinlets. Texture herbaceous or leathery, glabrous or pubescent, rarely scaly abaxially. Sori round or elongated, in one regular row on each side of costa or scattered on abaxial surface of pinnae, naked, superficial, without paraphyses. Sporangia long stalked, annulus with 1416 cells. Spores brown, ellipsoidal, surfaces verrucate[??verrucose] with often constricted, acuminate or echinate elements. x = 12, n = 36 [??just x for genus descr.].
About 20 species: tropical and subtropical Asia; 16 species in China (seven endemic).
1a...... Sori in one row on either side of costae.
2a...... Lateral pinnae distinctly stalked .................................................................................................. 1. A. tenuicauda
2b..... Lateral pinnae sessile.
3a...... Scales brown, lanceolate, gradually narrowed into acuminate apex.
4a. Sori rounded, always solitary ............................................................................................ 2. A. wallichiana
4b..... Sori elliptic or two adnate together ..................................................................................... 3. A. intermedia
3b..... Scales brown, dark brown, or black, ovate, abruptly narrowed into subulate apex.
5a. Rhizome concealed by overlapping brown scales; plants terrestrial ........................... 4. A. tatsienensis
5b..... Rhizome with whitish bloom visible between sparse, dark brown to black scales; plants epiphytic 5. A. nigropaleacea
1b..... Sori in two or more rows on either side of costae or irregularly scattered.
6a...... Rhizome 1012 mm in diam.; fronds abaxially usually with whitish bloom.
7a...... Lateral pinnae sessile ...................................................................................................................... 13. A. wardii
7b..... Lateral pinnae stalked.
8a. Lateral pinnae opposite ..................................................................................................... 14. A. medogensis
8b..... Lateral pinnae alternate.
9a...... Lamina abaxially with glandular hairs ................................................................................ 15. A. caudata
9b..... Lamina abaxially without glandular hairs ...................................................................... 16. A. salicifolia
6b..... Rhizome 36 mm in diam.; fronds abaxially green.
10a.... Pinna apex caudate.
11a.... Lateral pinnae 58 pairs, abaxially glabrous or pubescent .................................................... 8. A. elegans
11b... Lateral pinnae 14 pairs, abaxially pubescent or tomentose.
12a.... Rhizome sparsely scaly, surface with whitish bloom; scales brown ..................... 6. A. himalayensis
12b... Rhizome densely scaly, surface concealed; scales whitish ......................................... 7. A. tomentosa
10b... Pinna apex subacute to acuminate.
13a.... Plants terrestrial; pinnae 1015 cm, abaxially glabrous .......................................................... 10. A. mairei
13b... Plants epiphytic; pinnae 512 cm, abaxially glabrous, pubescent or tomentose.
14a.... Pinnae 812 pairs, ca. 5 cm, abaxially white tomentose ....................................... 8. A. notholaenoides
14b... Pinnae 58 pairs, 1012 cm, abaxially glabrous or pubescent.
15a.... Lamina 1520 cm wide, pinnae abaxially glabrous or sparsely pubescent; sori irregularly scattered in 2 or more ill-defined rows ............................................................................................................... 11. A. lehmanni
15b... Lamina 2530 cm wide, pinnae abaxially densely pubescent; sori in 4 or 5 rows, sometimes in confluent pairs ..................................................................................................................................... 12. A. lungtauensis
狭羽节肢蕨 xia yu jie zhi jue
Polypodium tenuicaudum Hooker, Sp. Fil. 5: 90. 1864; Arthromeris notabilis Ching; Pleopeltis juglandifolia (D. Don) T. Moore var. tenuicauda (Hooker) Beddome; Polypodium wallichianum Sprengel var. tenuicaudum (Hooker) Hooker[??not in Tropicos].
Rhizome 68 mm in diam., densely scaly throughout; scales brown, lanceolate, margin ciliate, apex acuminate. Stipe stramineous, 1525 cm, glabrous; lamina imparipinnate, 3545 Χ 1525 cm; pinnae 812 pairs, subopposite, stalked, oblique, ovate-lanceolate, 1525 Χ 24 cm, base cuneate, apex long acuminate; upper pinnae usually fertile and slightly narrower; lamina herbaceous, both surfaces glabrous, Sori round, 1.52 mm in diam., in a single row on each side of costa, medial or slightly nearer costa, solitary between lateral veins.
Epiphytic on tree trunks or epilithic; 12002800 m. Xizang, Yunnan [NE India, N Myanmar].
单行节肢蕨 dan hang jie zhi jue
Polypodium wallichianum Sprengel, Syst. Veg. ed. 16, 4: 53. 1827; Polypodium juglandifolium D. Don (Prodr. Fl. Nepal.: 3. 1825), not Willdenow (1810), [nor Humbolt & Bonpland??]; Arthromeris juglandifolia (D. Don) J. Smith; Pleopeltis capitellata (Mettenius) Beddome; P. juglandifolia (D. Don) T. Moore; Polypodium capitellatum Wallich ex Mettenius.
Rhizome 1015 mm in diam., densely scaly throughout; scales brown, lanceolate, 1015 Χ 23 mm, margin sparsely toothed, apex acuminate. Stipe stramineous, 1530 cm, glabrous; lamina imparipinnate, oblong in outline, 4070 Χ 3040 cm; pinnae usually 510 pairs, subopposite, sessile, oblique, ovate-lanceolate, 1020 Χ 24 cm, base rounded, margin entire or undulate, apex acuminate; lamina herbaceous or subleathery, both surfaces glabrous, Sori round, in one row on each side of costa, medial or slightly nearer costa, solitary between lateral veins.
Epiphytic on tree trunks or epilithic; 15002500 m. Guizhou, Sichuan, Xizang, Yunnan [Bhutan, N India, N Myanmar, Nepal, N Vietnam].
中间节肢蕨 zhong jian jie zhi jue
Rhizome ca. 10 mm in diam., densely scaly; scales brown, lanceolate, ca. 10 mm, margin entire, apex acuminate. Stipe stramineous, ca. 20 cm, glabrous; lamina imparipinnate, oblong or broadly lanceolate in outline, 3040 Χ 2025 cm; pinnae 68 pairs, subopposite, sessile, spreading, ovate-lanceolate, 1520 Χ 54 cm, fertile pinnae slightly contracted, base rounded, margin entire, apex acuminate; lamina herbaceous, abaxially pale or glaucous, adaxially green, both surfaces glabrous. Sori round or elliptic, in one row along each side of costa, medial, usually in confluent pairs.
* Epilithic; ca. 2000 m. Xizang.
康定节肢蕨 kang din jie zhi jue
Polypodium tatsienense Franchet & Bureau in H. Christ, Bull. Soc. Bot. France 52(Mιm. 1): 19. 1905.
Rhizome 47 mm in diam., densely scaly; scales brown, ca. 6 Χ 1.2 mm, margin irregularly toothed, apex long caudate. Stipe stramineous to castaneous, 2030 cm, glabrous; lamina imparipinnate, oblong, 3040 Χ 2025 cm, or simple and ovate-lanceolate; pinnae (1)38 pairs, subopposite, sessile, ascending/spreading[????], oblique, ovate-lanceolate, 1518 Χ 34 cm, base broadly cuneate to rounded or rarely subcordate, margin entire, apex caudate; terminal pinna larger, usually narrowed from base to apex; lamina herbaceous or subleathery, abaxially pale green or glaucous, adaxially green, both surfaces glabrous. Sori round, ca. 2 mm in diam., in one row on each side of costa, medial or slightly nearer to costa, solitary between lateral veins.
Terrestrial on dry slopes or epilithic; 10001600 m. Sichuan, Yunnan [India (Sikkim), Nepal, N Thailand].
The record of Arthromeris tatsienensis from Sikkim is based on the determination by Fraser-Jenkins (Taxon. Revis. Indian Subcont Pteridoph.: 59. 2008) of the only collection of Arthromeris lungtauensis var. sikkimensis S. R. Ghosh (nom. inval., location of type not indicated) as this species.
黑鳞节肢蕨 hei lin jie zhi jue
Rhizome ca. 8 mm in diam., covered with whitish bloom, sparsely scaly; scales nearly black or black, stiff, straight, broad at base, ca. 10 mm, margin irregularly toothed, apex long caudate. Stipe dark stramineous or brown, 2025 cm, glabrous; lamina imparipinnate, oblong, 5060 Χ 2530 cm; pinnae 5 or 6 pairs, opposite, subsessile, oblique, broadly lanceolate, 1520 Χ 34 cm, base rounded or subcordate, apex caudate-acute, terminal pinna stalked; lamina papery or herbaceous, abaxially glaucous, adaxially green, both surfaces glabrous, Sori round, ca. 2 mm in diam., in one row on each side of costa, medial or slightly close to costa, solitary between lateral veins.
* Epiphytic on tree trunks in evergreen broad-leaved forests; ca. 1800 m. Xizang.
琉璃节肢蕨 liu li jie zhi jue
Rhizome ca. 5 mm in diam., covered with whitish bloom, sparsely scaly; scales brown, broad at base, 34 mm, margin ciliate or toothed, apex long caudate. Stipe stramineous or dark stramineous, 1020 cm, glabrous; lamina imparipinnate, ovate in outline, 1540 Χ 1020 cm, or simple and ovate-lanceolate; pinnae 14 pairs, opposite, sessile, ± spreading, far apart, ovate or ovate-lanceolate, 815 Χ 35 cm, base rounded or subcordate, margin entire, broadly cartilaginous, apex caudate; lamina herbaceous or papery, both surfaces pubescent or abaxially tomentose and adaxially glabrous. Sori round, small, mostly in 3 or 4 rows along costa, nearer costa, in 2 series between lateral veins, pairs sometimes ± confluent.
Epiphytic on tree trunks or epilithic; 17002800 m. Sichuan, Xizang, Yunnan [Bhutan, NE India, N Myanmar, Nepal].
1a. Rhizome scales with ciliate margin; lamina with both surfaces pubescent ......................... 6a. var. himalayensis
1b. Rhizome
scales with toothed margin; lamina abaxially tomentose, adaxially glabrous
..................................................................................................................................................... 6b.
var. niphoboloides
琉璃节肢蕨 (原变种) liu li jie zhi jue (yuan bian zhong)
Polypodium himalayense Hooker, Sp. Fil. 5: 91. 1863; Pleopeltis himalayensis (Hooker) Beddome; Polypodium venustum Wallich ex C. B. Clarke (1880), not Desvaux (1811), nor Blume (1828).
Rhizome broad at base, ca. 4 mm in diam., margin ciliate. Stipe stramineous or dark stramineous, 1020 cm, glabrous; lamina 2040 Χ 1020 cm; pinnae 14 pairs, ovate or ovate-lanceolate, 1015 Χ 34 cm, base rounded or subcordate. Lamina herbaceous, both surfaces pubescent. Sori conspicuous.
Epiphytic on tree trunks; 17002800 m. Sichuan, Xizang, Yunnan [Bhutan, NE India, N Myanmar, Nepal].
灰茎节肢蕨 hui jing jie zhi jue
Polypodium venustum var. niphoboloides C. B. Clarke, Trans. Linn. Soc. London Bot. 1: 567. 1880.
Rhizome narrowly lanceolate, 34 mm in diam., margin toothed. Stipe dark stramineous, 515 cm, glabrous; lamina 1525 Χ 1015 cm; pinnae 1 or 2 pairs, ovate-lanceolate, 812 Χ 45 cm, base rounded. Lamina herbaceous or papery, abaxially tomentose, adaxially glabrous. Sori buried in tomentum, hardly visible.
Epilithic; 20002600 m. Yunnan [Bhutan].
厚毛节肢蕨 hou mao jie zhi jue
Rhizome ca. 4 mm in diam., densely scaly throughout; scales brown at center, with pale or whitish margin, lanceolate, broad at base, 67 mm, margin toothed, apex long caudate. Stipe dark stramineous, 1520 cm, glabrous; lamina imparipinnate, 2030 Χ 1015 cm, or simple and ovate to oblong; pinnae usually 1 or 2 pairs, opposite, sessile, ovate-lanceolate, 1012 Χ 34 cm, base rounded or cordate, overlapping rachis, margin entire, apex caudate or rarely laciniate; lamina herbaceous, abaxially tomentose, adaxially glabrous. Sori round, mostly in 3 or 4(or 5) rows along costa, in 2 series between lateral veins.
* Epiphytic on tree trunks; ca. 2600 m. Yunnan.
Fraser-Jenkins (Taxon. Revis. Indian Subcont. Pteridoph.: 60. 2008) believes that Arthromeris tomentosa is not separable from A. himalayensis.
[??vernacular name]
Epiphytic; ca. 1600 m. SE Xizang [India (N Arunachal Pradesh)]. [??what is latest FOC standard on this geopolitical issue]
美丽节肢蕨 mei li jie zhi jue
Rhizome 46 mm in diam., densely scaly; scales brown at center, with pale or whitish margin, ovate-lanceolate or lanceolate, broad at base, 812 mm, margin toothed or ciliate, apex acuminate. Stipe stramineous, 1525 cm, glabrous; lamina imparipinnate, 2535 Χ 1520 cm; pinnae 58 pairs, opposite, sessile, oblique or spreading, narrowly lanceolate to elliptic, 1216 Χ 1.52 cm, base rounded or cordate, cartilaginous margin narrow, apex long caudate; lamina herbaceous, both surfaces glabrous, or sometimes abaxially pubescent. Sori round, almost confluent when mature, in 2(or 3) rows along costa, closer to costa. in 2 series between lateral veins.
Epiphytic on tree trunks; 20002600(3400) m. ?Xizang, Yunnan [N Myanmar].
extended distribution fide FCCL
Two forms of Arthromeris elegans have been recognized: f. elegans with rhizome scales with toothed margins and with the lamina adaxially glabrous, and f. pianmaensis S. G. Lu (Acta Bot. Yunnan. 20: 406. 1998) with rhizome scales with ciliate margins and with the lamina adaxially pubescent.
多羽节肢蕨 duo yu jie zhi jue
Polypodium mairei Brause, Hedwigia 54: 208. 1913[??1914]; Arthromeris lehmannii (Mettenius) Ching var. auriculata Ching; A. tsayuensis Ching & Y. X. Lin; P. lehmanni (Mettenius) Ching var. mairei (Brause) C. Christensen.
Rhizome 56 mm in diam., densely scaly; scales light brown or glaucous, ovate-lanceolate, margin ciliate, apex acuminate. Stipe stramineous or castaneous, 1525 cm, glabrous; lamina imparipinnate, ovate-lanceolate, 3050 Χ 1525 cm; pinnae 812 pairs, opposite[????], sessile, ascending, ovate-lanceolate, 1015 Χ 23 cm, base rounded, margin entire or undulate, apex acuminate; lamina herbaceous, both surfaces glabrous, Sori round, [how large?], in 2 or 3 irregular rows along costae of distal pinnae, in 2 series between veins, pairs sometimes confluent.
Terrestrial on slopes in forests; 10002700 m. Guangxi, Guizhou, Hubei, Jiangxi, Shanxi, Sichuan, Xizang, Yunnan [N India, Myanmar].
Fraser-Jenkins (Taxon. Revis. Indian Subcont. Pteridoph.: 58. 2008) suggests that Arthromeris mairei might not be separable from A. moulmeinense Bedddome (1867).
节肢蕨 jie zhi jue
Polypodium lehmannii Mettenius, Abh. Senckenberg. Naturf. Ges. 117. 1856[??1857]; Arthromeris pinnata (Hayata) Ching; A. tibetana Ching; A. tibetana var. glabrescens Ching & S. K. Wu; Pleopeltis lehmanni (Mettenius) Beddome; Polypodium pinnatum Hayata; P. quasipinnatum Hayata.
Rhizome 45 mm in diam., densely or sparsely scaly and covered with whitish bloom; scales brown at center, paler toward margin, broad at peltate base, 46 mm, margin minutely toothed, apex long caudate. Stipe stramineous or light castaneous, 1020 cm, glabrous; rachis stramineous, glabrous; lamina imparipinnate, oblong-ovate in outline, 3040 Χ 1520 cm; pinnae usually 58 pairs, subopposite, sessile, spreading or slightly ascending, straight, lanceolate, base rounded or subcordate, overlapping rachis, margin entire or undulate with cartilaginous membrane up to 1.5 mm wide, apex acuminate. terminal pinna same as lateral pinnae; lamina papery or herbaceous, green, both surfaces usually glabrous, rarely abaxially sparsely pubescent. Sori round or elliptic, variable in size, mostly in 2 or 3 rows along costa, in 1 series between lateral veins.
Epiphytic on tree trunks or epilithic; 5002900 m. Guangxi, Guangdong, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, N India, Myanmar, Nepal, Thailand; Indochina to Philippines[please list specific countries]].
龙头节肢蕨 long tou jie zhi jue
Polypodium lungtauense (Ching) Ching ex C. Christensen[??ex or in].
Rhizome 45 mm in diam., densely scaly throughout; scales dark brown at center, paler toward margin, ovate-lanceolate, margin toothed, apex acuminate. Stipe dark stramineous or light castaneous, 1020 cm, glabrous; rachis and costae stramineous, pubescent; lamina imparipinnate, oblong or deltoid-lanceolate, 3040 Χ 2530 cm; pinnae 57 pairs, opposite[????], sessile, ascending/patent[????], lanceolate or ovate-lanceolate, 1012 Χ 23 cm, base rounded or cordate, cartilaginous margin entire, very narrow, apex acuminate; lamina papery, both surfaces densely pubescent. Sori round to elliptic, ca. 1 mm, mostly in 3 or 4 rows along costa, in 2 series between veins, pairs sometimes confluent.
Epiphytic on tree trunks or on rocks; 5002500 m. Fujian, Guangxi, Guangdong, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang [Laos, Nepal, Vietnam].
Material of Arthromeris lungtauensis has been incorrectly identified as Polypodium himalayense Hooker: H. Christ (Bull. Acad. Gιogr. Bot. 1902: 218. 1902; Wu, Wang & Pong, Bull. Dept. Biol. Sun Yatsen Univ. 3: 306. pl. 144. 1932).
灰背节肢蕨 hui bei jie zhi jue
Polypodium wardii C. B. Clarke, J. Linn. Soc., Bot. 25: 99. 1889; Arthromeris longipinna Ching & Y. X. Lin; Pleopeltis wardii (C. B. Clarke) Beddome.
Rhizome 1012 mm in diam., densely scaly throughout; scales brown, narrowly lanceolate, ovate at peltate base, 610 mm, margin subentire, apex acuminate. Stipe stramineous or dark stramineous, 3040 cm, glabrous; lamina imparipinnate, ovate-lanceolate, 4080(120) Χ 3050 cm; pinnae 58(16) pairs, subopposite, sessile, spreading or ascending, ovate-lanceolate, 1520(30) Χ 46(8) cm, base rounded or truncate, margin entire with [how broad???] cartilaginous membrane, apex caudate; lamina papery, abaxially usually glaucous on abaxial surface, adaxially green, both surfaces glabrous, Sori round, up to 2.5 mm in diam., mostly in 2(or 3) rows along costa in I[??1] series between veins, sometimes eventually almost confluent.
Epiphytic on tree trunks; 18002500 m. Xizang, Yunnan [Bhutan, N India, Myanmar, Nepal].
墨脱节肢蕨 mo tuo jie zhi jue
Rhizome ca. 6 mm in diam., densely scaly; scales brown, ovate at base, margin ciliate, apex long caudate. Stipe stramineous, 2030 cm, glabrous; lamina imparipinnate, 2030 Χ 1416 cm; pinnae usually 2 or 3 pairs, opposite, distinctly stalked, ascending, ovate or oblong, 1113 Χ 34 cm, base cuneate, margin entire with [how broad???] cartilaginous membrane, apex caudate; lamina papery, abaxially glaucous, adaxially green, both surfaces glabrous. Sori round, mostly in 3 or 4 rows along costa in 1 series between lateral veins.
* Epiphytic on tree trunks; ca. 1600 m. Xizang.
尾状节肢蕨 wei zhuang jie zhi jue
Rhizome ca. 10 mm in diam., densely scaly; scales brown, broad at base, margin ciliate, apex long caudate. Stipe stramineous at base, castaneous at upper part, 2535 cm, glabrous; lamina imparipinnate, 5060 Χ 2025 cm; pinnae usually 68 pairs, alternate, shortly stalked, ascending, oblong-lanceolate, 1822 Χ 45 cm, base cuneate, margin entire, broadly cartilaginous, apex caudate; lamina papery, abaxially glaucous with sparse glandular hairs, adaxially green. Sori round, mostly in 2 or 3 rows along costa in 1 series between lateral veins.
* Evergreen broad-leaved forests; ca. 1500 m. Xizang.
柳叶节肢蕨 liu ye jie zhi jue
Rhizome ca. 10 mm in diam., densely scaly; scales brown, ovate at base, 57 mm, margin shortly ciliate, apex long caudate. Stipe dark stramineous, 2530 cm, ca. 3 mm in diam., glabrous; lamina imparipinnate, 4050 Χ 1520 cm; pinnae usually 5 or 6 pairs, alternate, shortly stalked, ascending, ovate-lanceolate, 1520 Χ 35 cm, base attenuate into stalk, margin entire, with cartilaginous membrane, apex long caudate; lamina papery, abaxially light green or glaucous. adaxially dark green. Sori mostly round, ca. 1.5 mm diam., mostly in 2 rows along costa in 1 series between veins, rarely those nearest margin in confluent pairs.
* Epiphytic on tree trunks; 16001900 m. Xizang.
戟蕨属 ji jue shu
Ferns usually epiphytic, sometimes terrestrial, medium-sized. Rhizome long-creeping, densely covered with scales; scales dark brown, linear-lanceolate, peltate at base, entire or sparsely toothed at margin, opaque, hairlike at apex. Fronds remote, articulate to rhizome, dimorphic. Sterile fronds trilobate or pinnatifid, lobes broad, entire; veins reticulate with branched included veinlets; texture leathery, glabrous on both surfaces or scaly on abaxial surface when young, scales ovate, peltate. Fertile fronds strongly contracted, trilobate or pinnatifid; lobes linear-lanceolate. Sori covering whole abaxial surface of fertile fronds, mixed with short, simple or branched paraphyses. Sporangia with long stalks; annulus with ca. 14 indurate cells. Spores ellipsoidal, hyaline, shallowly tuberculate on surface. Chromosome number as yet unknown.
Three species: Tropical Asia; one species in China (Hainan).
Fraser-Jenkins (Taxon. Revis. Indian Subcont. Pteridoph.: 45. 2008) includes Christiopteris within the following genus, Selliguea. It is maintained by Christenhusz et al. (2011 [??citation]).
戟蕨 ji jue
Acrostichum tricuspe Hooker, Sp. Fil. 5: 272. 1864; Cheiropleuria tricuspis (Hooker) J. Smith; Gymnopteris tricuspis (Hooker) Beddome; Leptochilus tricuspis (Hooker) C. Christensen; Selliguea tricuspis (Hooker) Fraser-Jenkins.
Rhizome long-creeping, 56 mm in diam., densely covered with scales; scales reddish brown, ovate-lanceolate at base, 5-6 mm, minutely serrate at margin, subulate and long caudate at apex. Fronds remote. Stipe straw-colored or castaneous, 2035 cm, glabrous, stipes of fertile fronds longer than those of sterile fronds. Laminae dimorphic. Sterile lamina deeply 3-lobed or pinnatilobate, lobes 2035 Χ 2.57cm, central lobe larger than lateral lobes, entire at margin, acuminate at apex; veins reticulate with simple or branched included veinlets, hardly visible; texture leathery, pale green, glabrous on both surfaces. Fertile lamina with much narrower lobes, linear, 3040 Χ 11.5 cm. Sori with simple paraphyses.
Epiphytic on tree trunks or rarely terrestrial; 500800 m. Hainan [India (Sikkim), Malaysia, Thailand, Vietnam].
修蕨属 xiu jue shu
Crypsinus C. Presl; Phymatopsis J. Smith (1875), not Phymatopsis Tulasne ex Treviranus (1857); Phymatopteris Pichi Sermolli; Pichisermollia Fraser-Jenkins (2008), not [??H. C.] Monteiro Neto (1976 [??1980]); Pichisermollodes Fraser-Jenkins & Challis.
Rhizome slender, woody, densely scaly; scales reddish brown, brown, dark brown, or nearly black, rarely whitish, opaque, ovate-lanceolate or lanceolate, attenuate or setaceous, margin entire, toothed, or ciliate. Fronds remote, articulate to rhizome, monomorphic or dimorphic with longer and narrower fertile fronds. Stipe usually glabrous, rarely pubescent, scaly at base. Lamina simple, hastately 3-lobed, palmately or pinnately divided, sometimes pinnatisect with adnate pinnae, herbaceous or leathery, glabrous or pubescent, rarely scaly on abaxial side. Lobes usually lanceolate, with a distinct cartilaginous, entire, notched, or serrate margin, acuminate or obtuse at apex. Lateral veins (in lamina or segments) distinct; veinlets anastomosing to form areoles with included free veinlets. Sori round, in one row on either side of costa, less often linear or in many rows, usually superficial, rarely sunken on abaxial surface and raised on adaxial surface. Sporangia with long stalk, annulus with 14 indurate cells. Spores brown, ellipsoidal, surface tuberculate or verrucate. x = 12, n = 36, 37, 2n = 72. [??just x for genus description]
About 75 species: tropical and subtropical Asia, Australia, Pacific islands, South Africa, and Madagascar; 48 species in China (30 endemic).
Until relatively recently, most species of Selliguea were included within Phymatopteris but Smith et al. (2006[??citation]), Fraser-Jenkins (2008[??citation]) and Christenhusz et al. (2011[??citation]) all included Phymatopteris along with Crypsinus within Selliguea.
Fraser-Jenkins erected the genus Pichisermollodes for those species with Polypodium-like pinnatifid to pinnate fronds with notched and toothed margins but this genus was not accepted by Christenhusz et al.
1a...... All fronds simple, linear, lanceolate, or ovate.
2a...... Sori linear, between and parallel to adjacent lateral veins .................................................................... 48. S. feei
2b..... Sori round or less often elliptic, very rarely confluent.
3a...... Fronds moderately dimorphic; sterile lamina ovate, fertile lamina narrower, linear or ovate-lanceolate.
4a. Soriferous portion not or slightly contracted; sori round, separate ........................ 1. S. rhynchophylla
4b..... Soriferous portion much contracted, linear; sori elongate or confluent in upper part of frond 2. S. wuliangshanense
3b..... Fronds monomorphic; lamina oblong, ovate, or linear.
5a. Margin notched.
6a...... Sori sunken on abaxial surface and raised on adaxial surface ............................... 9. S. yakushimense
6b..... Sori superficial.
7a...... Lamina apex acuminate, base cuneate.
8a...... Lamina thin, pellucid; veinlets distinct ........................................................... 10. S. pellucidifolia
8b..... Lamina thick, opaque; veinlets obscure ..................................................................... 11. S. engleri
7b..... Lamina apex obtuse, base broadly cuneate to rounded.
9a...... Lamina herbaceous, stipe slender ............................................................................. 12. S. tenuipes
9b..... Lamina leathery, stipe thick ..................................................................................... 13. S. omeiensis
5b..... Margin entire.
10a.... Lamina apex rounded, obtuse, or acute.
11a.... Lamina apex rounded to obtuse; sori close to midrib ..................................................... 4. S. obtusa
11b... Lamina apex obtuse to acute; sori medial ............................................................... 5. S. oblongifolia
10b... Lamina apex acuminate or caudate.
12a.... Rhizome with whitish bloom and sparse scales; lamina margin wide, flat, translucent 3. S. chrysotricha
12b... Rhizome densely scaly; lamina margin narrow, thickened, not translucent.
13a.... Lamina base gradually narrowed; sori medial ..................................................... 6. S. hainanense
13b... Lamina base truncate, rounded, or broadly cuneate, sori close to midrib.
14a.... Lamina base truncate or rounded, abaxial surface glaucous ........................... 7. S. majoensis
14b... Lamina base broadly cuneate, abaxial surface yellowish green ................... 8. S. griffithiana
1b..... At least some fronds deeply lobed, hastately divided, palmately parted, pinnately parted, or pinnatisect.
15a.... Divided fronds at most hastate, with two basal lateral lobes.
16a.... Lamina margin entire or remotely notched; sori slightly sunken abaxially; scales yellowish brown 14. S. taiwanensis
16b... Lamina margin regularly notched; sori superficial; scales reddish brown ............................. 15. S. hastata
15b... At least some divided fronds palmate with more than two lateral lobes or pinnately divided.
17a.... Fronds palmately divided with 46 lobes (S. dactylina s.l.).
18a.... Stipe
castaneous; [frond/pinnae] lobes obtuse at apex; sori close to margin;
plant 510 cm high
....................................................................................................................................................... 16.
S. digitata
18b... Stipe straw-colored; [frond/pinnae] lobes acuminate at apex; sori medial; plant 2030 cm high.
19a.... Scales narrowly lanceolate, yellowish brown, 57 mm, gradually narrowed from peltate base to acuminate apex 17. S. dactylina
19b... Scales ovate, dark brown, ca. 10 mm, gradually or sometimes abruptly narrowed into long hairlike apex 18. S. chenopus
17b... Fronds pinnately divided, lateral lobes/pinnae 110 pairs.
20a.... Lamina margin entire.
21a.... Fronds pubescent; lateral lobes 13 pairs, apical lobes distinctly larger than lateral lobes 19. S. trisecta
21b... Fronds glabrous; lateral lobes 15(8) pairs, apical lobes similar to lateral lobes.
22a.... Fronds pinnatifid at base, lobes triangular or broadly lanceolate ............................ 20. S. oxyloba
22b... Fronds pinnatisect at base, pinnae ovate-lanceolate .......................................... 21. S. likiangensis
20b... Lamina margin notched or toothed.
23a.... Lamina margin with notches regularly placed between veins.
24a.... Fronds distinctly dimorphic; sori deeply sunken ........................................................ 22. S. trilobus
24b... Fronds monomorphic; sori superficial.
25a.... Lateral lobes 1 or 2 pairs, deltoid or subdeltoid; basal lobes spreading in trilobed lamina 23. S. cruciformis
25b... Lateral lobes 28 pairs, lanceolate; basal lobes ascending or deflexed.
26a.... Lateral lobes all ascending, lamina cuneate or rounded at base.
27a.... Fronds hairy throughout; terrestrial ................................................................. 24. S. hirtella
27b... Fronds glabrous; epiphytic.
28a.... Fronds 2550 cm; stipe usually longer than lamina .................................. 25. S. taeniata
28b... Fronds 1020 cm; stipe shorter than lamina.
29a.... Scales black with whitish cilia at margin ...................................... 26. S. kingpingensis
29b... Scales yellowish brown or whitish.
30a.... Scales
yellowish brown with entire margin; sori medial
.................................................................................................................. 27.
S. connexa
30b... Scales
whitish with ciliate margin; sori close to margin
..................................................................................................... 28.
S. daweishanensis
26b... Lowest lobes deflexed or slightly deflexed, lamina cordate or truncate at base.
31a.... Scales ovate, castaneous to black; abaxial side of lamina usually sparsely scaly especially on midrib and veins .................................................................................................................... 29. S. ebenipes
31b... Scales lanceolate, brown to whitish; abaxial side of lamina without scales.
32a.... Scales brown or dark brown.
33a.... Rhizome 812 mm in diam.; scales margin entire; sori sunken and close to margin 30. S. echinospora
33b... Rhizome 34 mm diam.; scales ciliate at margin; sori superficial, ± close to costa 31. S. quasidivaricata
32b... Scales with broad whitish or light brown margin, or dark brown when older.
34a.... Lamina glabrous on both surfaces; lobes acuminate at apex .............. 32. S. albopes
34b... Lamina pubescent on both surfaces; lobes obtuse at apex ......... 33. S. pianmaensis
23b... Lamina margin usually with a sharp tooth below each notch, often with additional teeth between each pair of notches.
35a.... Fronds pubescent on one or both surfaces.
36a.... Lobes ca. 1 cm wide, serrulate at margin ............................................................. 34. S. nigrovenia
36b... Lobes 1.52 cm wide, crenate at margin ......................................................... 35. S. incisocrenata
35b... Fronds glabrous on both surfaces.
37a.... Lateral lobes obtuse or acute at apex.
38a.... Lamina
58 cm; stipe straw-colored or dark straw-colored at base
................................................................................................................................ 36.
S. shensiense
38b... Lamina 520 cm; stipe purple or castaneous at base.
39a.... Lobes shallowly crenate to pinnately lobed ..................................... 37. S. crenatopinnata
39b... Lobes densely serrulate or duplicate-serrate at margin.
40a.... Lateral lobes ovate-lanceolate, constricted at base ............................. 38. S. glaucopsis
40b... Lateral lobes linear ........................................................................................ 39. S. conmixta
37b... Lateral lobes acuminate or caudate-acuminate at apex.
41a.... Lowest lobes of lamina deflexed.
42a.... Lateral lobes gradually narrowed from base to apex ................................... 40. S. stracheyi
42b... Lateral lobes ovate-lanceolate, slightly constricted at base ..................... 41. S. conjuncta
41b... Lowest lobes of lamina ascending.
43a.... Scales black without light margin but with whitish cilia on margin.
44a.... Lobes obtusely serrate at margin ................................................................ 42. S. stewartii
44b... Lobes sharply serrate at margin.
45a.... Lamina with 49 pairs of lateral pinnae, cartilaginous margin straw-colored 43. S. nigropaleacea
45b... Lamina
with 2 or 3 pairs of lateral lobes, cartilaginous margin reddish
......................................................................................................... 44.
S. roseomarginata
43b... Scales brown or dark brown or with a black center, with distinct, lighter, dentate or ciliate margin.
46a.... Scales dark brown or castaneous; lateral veins ending in margin .......... 45. S. tibetana
46b... Scales brown; lateral veins not ending in margin.
47a.... Marginal teeth aristate ......................................................................... 46. S. malacodon
47b... Marginal teeth not aristate ............................................................... 47. S albidoglauca
喙叶假瘤蕨 hui ye jia liu jue
Polypodium rhynchophyllum Hooker, Hookers Icon. Pl. 6: t. 954. 1854; Crypsinus okamotoi (Tagawa) Tagawa; C. rhynchophyllus (Hooker) Copeland; Phymatodes okamotoi Tagawa; P. rhynchophylla (Hooker) Ching; Phymatopsis rhynchophylla (Hooker) J. Smith; Phymatopteris rhynchophylla (Hooker) Pichi-Sermolli; Pleopeltis rhynchophylla (Hooker) T. Moore; Selliguea okamotoi (Tagawa) Ralf Knapp.
Rhizome 12 mm in diam., densely scaly throughout; scales brown, ovate-lanceolate, ca. 5 mm, margin entire to sparsely toothed, apex acuminate. Fronds dimorphic. Sterile fronds: stipe 12 cm; lamina simple, ovate or oblong, 15 Χ 12 cm, base cordate, margin notched, apex obtuse. Lateral veins distinct. Lamina herbaceous or subleathery, both surfaces glabrous, abaxial surface usually purplish, adaxial surface green. Fertile fronds: stipe 510 cm; lamina linear or lanceolate, 520 Χ 12 cm, soriferous portion not or only slightly contracted. Sori round, medial.
Epiphytic on tree trunks; 12002700 m. Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan [Cambodia, N India, Indonesia, Laos, Myanmar, Nepal, Philippines, Thailand, Vietnam].
Polypodium drymoglossoides H. Christ (1904) not Baker (1887): ?misidentification, needs checking.
无量山假瘤蕨 wu liang shan jia liu jue
Phymatopteris wuliangshanensis W. M. Chu, Acta Bot. Yunnan., Suppl. 5: 58. 1992; Crypsinus wuliangshanensis (W. M. Chu) X. Cheng.
Rhizome 12 mm in diam., densely scaly; scales pale brown, lanceolate, 34 Χ 11.5 mm, margin entire, apex acuminate. Fronds dimorphic. Sterile fronds: stipe 1.52 cm; lamina simple, circular or ovate, 23 Χ 12 cm, base cordate, margin notched, apex rounded. Lateral veins visible, veinlets obscure. Lamina herbaceous or subleathery, both surfaces glabrous, pale green, Fertile fronds: stipe stramineous, 25 cm, slender, glabrous; lamina lower portion ovate-lanceolate, 510 Χ 12 cm, base cuneate, soriferous portion contracted. Sori separate at base of soriferous part, forming a coenosorus toward apex.
* Epiphytic on tree trunks, evergreen broad-leaved forests; 22002500 m. Yunnan (Wuliang Shan).
白茎假瘤蕨 bai jing jia liu jue
Polypodium chrysotrichum C. Christensen, Contr. U.S. Natl. Herb. 26: 320. 1931; Crypsinus chrysotrichus (C. Christensen) Tagawa; Phymatodes chrysotricha (C. Christensen) Ching; Phymatopsis chrysotricha (C. Christensen) Ching; Phymatopteris chrysotricha (C. Christensen) Pichi Sermolli.
Rhizome with whitish bloom, sparsely scaly; scales brown, ovate at peltate base, margin entire, apex acuminate. Fronds ± monomorphic (fertile fronds slightly narrower and more oblong). Stipe straw-colored, 510 cm, glabrous; lamina simple, ovate to oblong, 510 Χ 36 cm, base rounded to shallowly cordate, margin entire, wide, flat, translucent, apex caudate. Veins all distinct on both surfaces. Lamina leathery, both surfaces glabrous, abaxial surface pale green, adaxial surface green and shiny. Sori round, medial.
* Epiphytic on tree trunks; 22002900 m. Yunnan [Myanmar].
圆顶假瘤蕨 yuan din jia liu jue
Phymatopsis obtusa Ching, Acta Phytotax. Sin. 9: 184. 1964; Phymatopteris obtusa (Ching) Pichi Sermolli.
Rhizome ca. 3 mm in diam., densely scaly; scales brown, lanceolate, ca. 5 mm, margin entire, apex acuminate. Stipe straw-colored or light brown, 610 cm, glabrous; lamina simple, oblong or ovate, 515 Χ 23 cm, base cordate, margin entire or undulate, apex rounded or obtuse. Lateral veins thick, distinct, ascending, veinlets obscure. Lamina leathery, both surfaces glabrous. Sori round, close to midrib.
* Epiphytic on tree trunks or on rocks; 14001700 m. Guangxi, Hainan, Xizang, Yunnan.
长圆假瘤蕨 chang yuan jia liu jue
Phymatopsis oblongifolia S. K. Wu, Acta Phytotax. Sin. 23: 400. 1985; Crypsinus oblongifolius (S. K. Wu) X. Cheng; Phymatopteris oblongifolia (S. K. Wu) W. M. Chu & S. G. Lu.
Rhizome ca. 3 mm in diam., densely scaly throughout; scales reddish brown, broad at peltate base, margin sparsely toothed, apex acuminate. Fronds monomorphic. Stipe straw-colored, 515 cm, ca. 1 mm in diam., glabrous; lamina simple, oblong or ovate, 515 Χ 23 cm, base cordate, margin entire, apex obtuse or acute. Lateral veins distinct on both surfaces, veinlets obscure. Lamina leathery, both surfaces glabrous. Sori round, slightly near midrib.
* Epiphytic on tree trunks; ca. 1400 m. Yunnan.
海南假瘤蕨 hai nan jia liu jue
Phymatodes hainanensis Ching, Contr. Inst. Bot. Natl. Acad. Peiping 2(3): 68. 1933; Crypsinus hainanensis (Ching) Tagawa; Phymatopsis cunea Ching; P. hainanensis (Ching) Ching; Phymatopteris cunea (Ching) Pichi Sermolli; P. hainanensis (Ching) Pichi Sermolli; Polypodium echinosporum C. Christensen.
Rhizome ca. 3 mm in diam., densely scaly; scales dark brown in center, paler toward margins, ovate at peltate base, ca. 3 mm, margin entire, apex subulate. Fronds monomorphic. Stipe straw-colored, 813 cm, ca. 1 mm in diam., glabrous; lamina simple, ovate-lanceolate, 815 Χ 1.53 cm, widest around middle and gradually narrowed toward both ends, base cuneate, margin entire, apex acuminate. Lateral veins visible, veinlets obscure. Lamina papery, both surfaces glabrous. Sori round, slightly near midrib and slightly raised on adaxial surface.
* Epiphytic on tree trunks or on rocks in forests; 500600 m. Hainan, Yunnan.
宽底假瘤蕨 kuan di jia liu jue
Polypodium majoense C. Christensen in H. Lιveillι, Cat. Pl. Yun-Nan: 108. 1916; Crypsinus majoensis (C. Christensen) X. Cheng; Phymatodes griffithiana (Hooker) Ching var. majoensis (C. Christensen) Ching; Phymatopsis majoensis (C. Christensen) Ching; Phymatopteris majoensis (C. Christensen) Pichi Sermolli.
Rhizome 34 mm in diam., densely scaly; scales brown, lanceolate, 45 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored, 1015 cm, glabrous; lamina simple, oblong, 1525 Χ 36 cm, broadest near base, base rounded or truncate, margin entire, apex shortly acuminate. Lateral veins visible, veinlets obscure. Lamina subleathery, both surfaces glabrous, abaxial surface glaucous, adaxial surface pale green. Sori round, close to midrib.
* Epiphytic on tree trunks or on rocks; 14001800 m. Anhui, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Yunnan.
Hovenkamp believes that Selliguea majoensis would be better treated as a variety of the following species, S. griffithiana (Hooker) Fraser-Jenkins.
大果假瘤蕨 da guo jia liu jue
Polypodium griffithianum Hooker, Hookers Icon. Pl. 10[??6]: t. 951. 1854; Crypsinus griffithianus (Hooker) Copeland; Phymatodes griffithiana (Hooker) Ching; Phymatopsis griffithiana (Hooker) J. Smith; P. integerrima Ching; Phymatopteris griffithiana (Hooker) Pichi Sermolli; P. integerrima (Ching) Bir; Pleopeltis griffithiana (Hooker) T. Moore.
Rhizome 34 mm in diam., densely scaly; scales brown, lanceolate, ca. 5 mm, margin entire, apex long acuminate. Fronds monomorphic. Stipe straw-colored, 515 cm, densely scaly at base, subglabrous or with hairlike scales upward; lamina simple, oblong or ovate-lanceolate, 1025 Χ 34 cm, usually widest below middle, broadly cuneate at base, margin entire or shallowly undulate, usually revolute, apex shortly acuminate. Lateral veins distinct on both surfaces, veinlets obscure. Lamina leathery or subleathery, both surfaces glabrous, abaxial surface pale green, adaxial surface green. Sori round, large, close or slightly close to midrib.
Epiphytic on tree trunks or on rocks; 13003200 m. Anhui, Guizhou, Sichuan, Xizang, Yunnan [Bhutan, N India, Myanmar, Nepal, Thailand, Vietnam].
屋久假瘤蕨 wu jiu jia liu jue
Polypodium engleri Luerssen var. yakushimense Makino, Bot. Mag. (Tokyo) 23: 248. 1909; Crypsinus yakushimensis (Makino) Tagawa; Phymatodes yakushimensis (Makino) Tagawa; Phymatopsis fukienensis Ching; P. yakushimensis (Makino) H. Itτ; Phymatopteris fukienensis (Ching) Pichi Sermolli; P. yakushimensis (Makino) Pichi Sermolli; P.[??genus] yakushimense (Makino) Makino & Nemoto.
Rhizome ca. 3 mm in diam., densely scaly; scales brown, lanceolate, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored, 515 cm, slender, glabrous; lamina simple, linear-elliptic, 515 Χ 12 cm, base cuneate, margin notched, apex acuminate. Lateral veins distinct, veinlets obscure. Lamina papery, both surfaces glabrous, abaxial surface glaucous, adaxial surface green. Sori round, medial, sunken on abaxial surface and raised on adaxial surface.
On muddy rocks along streams, usually rheophytic in flood zone, in forest understories; 200800 m. Fujian, Guangxi, Guizhou, Hunan, Jiangxi, Taiwan, Zhejiang [Japan, Korea].
透明叶假瘤蕨 tou ming ye jia liu jue
Polypodium pellucidifolium Hayata, Icon. Pl. Formosan. 4: 250. 1914; Phymatopsis pellucidifolia (Hayata) H. Itτ; Phymatopteris pellucidifolia (Hayata) Pichi Sermolli.
Rhizome 34 mm in diam., densely scaly; scales ovate-lanceolate, rounded at base, margin entire, apex acuminate. Stipe light brown, 910 cm, glabrous; lamina simple, linear or oblanceolate, 3035 Χ 22.2 cm, base cordate, margin entire or undulate, apex shortly acuminate. Veins all distinct. Lamina membranous, pellucid, both surfaces glabrous. Sori round, small, slightly close to midrib.
* On rocks beside streams of forest understories. Taiwan (Alishan).
恩氏假瘤蕨 en shi jia liu jue
Polypodium engleri Luerssen, Bot. Jahrb. Syst. 4(4): 361. 1883; Crypsinus engleri (Luerssen) Copeland; Crypsinus engleri var. coriaceus (Tagawa) Tagawa; Phymatodes engleri (Luerssen) Ching; Phymatopsis engleri (Luerssen) H. Itτ; Phymatopteris engleri (Luerssen) Pichi Sermolli; P. engleri var. coriacea Tagawa[??not in Tropicos]; Polypodium engleri var. hypoleuca Hayata; P. hastatum Thunberg var. engleri (Luerssen) H. Christ.
Rhizome 34 mm in diam., densely scaly; scales reddish brown, linear to linear-lanceolate, 36 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored or light brown, 515(20) cm, glabrous; lamina simple, linear or oblanceolate, 515(28) Χ 13 cm, widest above middle, base cuneate, margin entire or slightly crenate, apex shortly acuminate. Lateral veins visible, veinlets obscure. Lamina papery or subleathery, stiff, both surfaces glabrous, abaxial surface usually glaucous. Sori round, small, slightly closer to midrib, superficial.
Epiphytic on tree trunks or on rocks. Fujian, Guangxi, Guizhou, Jiangxi, Taiwan, Zhejiang [Japan, Korea].
细柄假瘤蕨 xi bing jia liu jue
Phymatopsis tenuipes Ching, Acta Phytotax. Sin. 9: 187. 1964; Phymatopteris tenuipes (Ching) Pichi Sermolli.
Rhizome ca. 2 mm in diam., densely scaly; scales yellowish brown, lanceolate, 34 mm, margin entire, apex acuminate. Stipe straw-colored, 24 cm, slender, glabrous; lamina simple, narrowly oblong or obovate, 37 cm Χ 57 mm, base rounded or broadly cuneate, margin notched, apex obtuse. Lateral veins visible, veinlets obscure. Lamina herbaceous, both surfaces glabrous, abaxial surface glaucous, adaxial surface green. Sori round, small, medial, borne on upper portion of lamina.
* Epiphytic on tree trunks or on rocks; 13001700 m. Guizhou, Sichuan.
峨嵋假瘤蕨 e mei jia liu jue
Phymatopsis omeiensis Ching, Acta Phytotax. Sin. 9: 187. 1964; Phymatopteris omeiensis (Ching) Pichi Sermolli.
Rhizome ca. 2 mm in diam., densely scaly; scales brown, lanceolate, 35 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored, 35 cm, glabrous; lamina simple, ovate-lanceolate, 812 Χ 12 cm, lower part widest and sterile, base broadly cuneate or rounded, margin notched, apex acute or obtuse. upper part soriferous, slightly contracted, Lateral veins visible, veinlets obscure. Lamina leathery, both surfaces glabrous, pale green. Sori round, slightly close to midrib.
* Epiphytic on rocks. Sichuan (Emei Shan).
台湾假瘤蕨 tai wan jia liu jue
Phymatodes taiwanensis Tagawa, Acta Phytotax. Geobot. 11: 310. 1942; Crypsinus taiwanensis (Tagawa) Tagawa; Phymatopsis taiwanensis (Tagawa) Ching; Phymatopteris taiwanensis (Tagawa) Pichi Sermolli.
Rhizome 23 mm in diam., densely scaly; scales yellowish brown, lanceolate, broad at peltate base, apex long acuminate with narrow subulate tip. Stipe straw-colored, 816 cm, glabrous; lamina simple or trifid within one individual; simple lamina linear, 816 Χ ca. 1 cm, base cordate, margin entire or remotely notched, apex caudate; trifid lamina: central lobe much longer than lateral lobes, 1624 Χ ca. 1 cm, all lobes usually slightly contracted at base, widest at middle, apex acuminate. Lateral veins visible, veinlets obscure. Sori round, slight sunken on abaxial surface and raised on adaxial surface.
* On rocks, [??valleys] between mountains; 15002000 m. Taiwan.
金鸡脚假瘤蕨 jin ji jiao jia liu jue
Polypodium hastatum Thunberg, Syst. Veg., ed. 14 (J. A. Murray),: 335[??935]. 1784; Crypsinus hastata (Thunberg) Copeland; Drynaria hastata (Thunberg) Fιe; Phymatodes hastata (Thunberg) Ching; Phymatopsis chenkouensis Ching; P. hastata (Thunberg) Kitagawa ex H. Itτ; P. hastata f. arenaria (Baker) Ching; P. hastata f. dolichopoda (Diels) Ching; P. hastata f. nikkoensis (H. Christ ex Matsumura) H. Itτ; P. hastata f. pygmaea (Maximowicz) H. Itτ; P. hastata var. pygmaea (Maximowicz) Li & J. Z. Wang; P. hastata f. simplex (H. Christ) Ching; P. hunyaensis Ching; P. rotunda Ching; P. shandongensis J. X. Li & C. Y. Wang; P. similis (Ching) C. Y. Wu[??Ching]; P. simplicifolia Ching; P. tarningensis Ching; Phymatopteris chenkouensis (Ching) Pichi Sermolli; P. hastata (Thunberg) Pichi Sermolli; P. hunyaensis (Ching) Pichi-Sermolli; P. rotunda (Ching) Pichi Sermolli; P. simplicifolia (Ching) Pichi Sermolli; Pleopeltis hastata (Thunberg) T. Moore; Polypodium arenarium Baker; P. dolichopodum Diels; P. hastatum var. dolichopodum (Diels) C. Christensen; P. hastatum var. nikkoense H. Christ ex Matsumura; P. hastatum f. pygmaeum Maximowicz; P. hastatum var. simplex H. Christ.
Rhizome 23 mm in diam., densely scaly throughout; scales reddish brown, lanceolate, ca. 5 mm, broad at peltate base, margin entire or sometimes sparsely toothed, apex long acuminate. Fronds monomorphic. Stipe straw-colored, 220 cm, 12 mm in diam., glabrous; lamina simple or hastate; simple lamina: ovate to linear or lanceolate, 220 Χ 12 cm, base cuneate or rounded, margin notched, apex acute or obtuse; hastate lamina: with one or two basal lateral lobes, sometimes one much longer than other, all lobes widest at base, apices acuminate. Costa and lateral veins distinct, veinlets obscure. Lamina papery or herbaceous, both surfaces glabrous, abaxial surface usually somewhat glaucous. Sori round, large, 23 mm in diam., medial, superficial.
Terrestrial along paths on slopes, common. Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shandong, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Japan, Korea, Philippines, Russia].
掌叶假瘤蕨 zhang ye jia liu jue
Phymatodes digitata Ching, Contr. Inst. Bot. Natl. Acad. Peiping 2(3): 77. 1933; Crypsinus digitatus (Ching) Tagawa; Phymatopsis digitata (Ching) Ching; P. palmatifida Ching & K. H. Shing[??Ching & Chiu]; Phymatopteris digitata (Ching) Pichi Sermolli; Polypodium koi C. Christensen.
Rhizome ca. 4 mm in diam., densely scaly throughout; scales light brown, lanceolate, ca. 5 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe castaneous, 210 cm, scaly at base, glabrous upward; lamina palmately 25-lobed, 59 Χ 59 cm, base rounded. margin entire or undulate. Lobes linear, central lobe usually longer than lateral lobes, 510 Χ 11.2 cm, apex obtuse or acute. Costa distinct, lateral veins and veinlets obscure. Lamina herbaceous, both surfaces glabrous, abaxial surface glaucous, adaxial surface green. Sori round, close to margin, superficial.
* Epiphytic on tree trunks on hill tops; 10001400 m. Guangdong, Guizhou, Zhejiang.
指叶假瘤蕨 zhi ye jia liu jue
Polypodium dactylinum H. Christ, Bull. Soc. Bot. France 52(Mιm. 1): 20. 1905; Crypsinus dactylinus (H. Christ) Tagawa; Phymatodes dactylina (H. Christ) Ching; Phymatopsis dactylina (H. Christ) Ching; Phymatopteris dactylina (H. Christ) Pichi Sermolli.
Rhizome 35 mm in diam., densely scaly throughout; scales yellowish brown, narrowly lanceolate, 57 mm, margin entire, apex long acuminate or setaceous. Fronds monomorphic. Stipe straw-colored, 710 cm, glabrous; lamina palmately 47-lobed, 1020 Χ 1015 cm, base cuneate or cordate, margin entire and slightly revolute, Lobes narrowly oblong, central lobe longer than lateral lobes, 510 Χ 11.5 cm, outermost lobes much shorter, apex obtuse or acute. Costa distinct, lateral veins and veinlets obscure. Lamina herbaceous, both surfaces glabrous, abaxial surface pale green, grayish when dried, adaxial surface green. Sori round, medial or slightly close to margin.
* Epiphytic on tree trunks or on rocks; 12001400 m. Sichuan, Zhejiang.
鹅绒假瘤蕨 e rong jia liu jue
Polypodium chenopus H. Christ, Bull. Soc. Bot. France 52(Mιm. 1): 20. 1905; Phymatopteris chenopus (H. Christ) S. G. Lu.
Rhizome 45 mm in diam., densely scaly throughout; scales dark brown or nearly black, gradually or abruptly narrowed upward, ca. 10 mm, broad at peltate base, margin setaceous from middle to apex. Fronds monomorphic. Stipe straw-colored, 1015 cm, glabrous; lamina palmately 46-lobed, 1020 Χ 1015 cm, base cuneate or cordate. margin entire and slightly revolute. Lobes linear, central lobe longer than lateral lobes, 1020 Χ 12 cm, apices acute or obtuse. Costa distinct, lateral veins and veinlets obscure. Lamina herbaceous, both surfaces glabrous, abaxial surface glaucous, adaxial surface green. Sori round, medial or slightly close to margin.
* Epiphytic on tree trunks or on rocks; 18003400 m. Yunnan (Deqin, Gongshan, Lijiang).
三出假瘤蕨 san chu jia liu jue
Polypodium trisectum Baker, Bull. Misc. Inform. Kew 1898: 232. 1898; Crypsinus hirsutus Tagawa & K. Iwatsuki; C. trisectus (Baker) Tagawa; Phymatodes trisecta (Baker) Ching; Phymatopsis trisecta (Baker) Ching; Phymatopteris trisecta (Baker) Pichi Sermolli; P. trisecta var. hirticarpa Ching[??not in Tropicos]; Polypodium podobasis H. Christ.
Rhizome 34 mm in diam., densely scaly throughout; scales dark brown at center, paler toward margin, ovate-lanceolate, 45 Χ 11.5 mm, margin ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 612 cm, hairy throughout; lamina hastate to pinnatifid, 2025 Χ 1520 cm, base broadly cuneate or cordate, margins entire or undulate. Lateral lobes 13 pairs, ascending, broadly lanceolate, gradually narrowed from base to acuminate apex, apical lobe longer than lateral ones, 1020 Χ 24 cm, ± contracted at base, apex acuminate, Costa raised on both surfaces, lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces densely pubescent. Sori round, large, medial or slightly close to costa.
Terrestrial in forests; 16002400 m. Sichuan, Yunnan [Myanmar, Thailand].
尖裂假瘤蕨 jian lie jia liu jue
Polypodium oxylobum Wallich ex Kunze, Linnaea 24: 255. 1851; Crypsinus kwangtungensis (Ching) Tagawa; C. oxylobus (Wallich ex Kunze) Sledge; C. pingpienensis (Ching) Nakaike [pingpiensis]; Phymatodes kwangtungensis Ching; P. oxyloba (Wallich ex Kunze) C. Presl ex Ching[??okay]; Phymatopsis kwangtungensis (Ching) Ching; P. pingpienensis Ching; P. suboxyloba Ching; P. trifida (D. Don) J. Smith; Phymatopteris kwangtungensis (Ching) Pichi Sermolli; P. oxyloba (Wallich ex Kunze) Pichi Sermolli; P. pingpienensis (Ching) Pichi Sermolli; P. suboxyloba (Ching) Pichi Sermolli; Pleopeltis oxyloba (Wallich ex Kunze) Beddome; P. trifida (D. Don) Beddome; Pleuridium oxyloba (Wallich ex Kunze) J. Smith; Polypodium hastatum var. oxylobum (Wallich ex Kunze) C. B. Clarke; P. kwangtungense (Ching) Ching ex C. Christensen[??authors]; P. longipes Ching (1931), not Link ex Kuntze (1850), nor Fιe (1872); P. oxylobum (Wallich ex Kunze) Ching [??authors]; P. trifidum D. Don (1825), not Wither. (1796), nor Hoffm. (1790).
Rhizome 45 mm in diam., densely scaly throughout; scales brown at center, paler toward margin, lanceolate, ca. 5 mm, margin and sometimes surface ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored or light brown, usually 1020 cm, densely scaly at base, glabrous upward; lamina trifid or pinnatifid, incised up to 515 mm from costa, 2030 Χ 1020 cm, base broadly cuneate. margin entire or undulate. Lateral lobes (1 or)25(8) pairs, ascending, deltoid or broadly lanceolate, 1015 Χ 1.55 cm, base sometimes slightly contracted, apex acuminate. Costa and lateral veins raised on both surfaces. Lamina papery, both surfaces glabrous. Sori round, large, slightly closer to costa.
Epiphytic on rocks or on tree trunks in evergreen forests, or terrestrial; 10002700 m. Guangdong, Guangxi, Sichuan, Yunnan [N India, Myanmar, Nepal, Thailand, Vietnam].
The name Polypodium oxylobum was first introduced by Wallich (Numer. List no. 294. 1828, nom. nud.).
丽江假瘤蕨 li jiang jia liu jue
Phymatopsis likiangensis Ching, Acta Phytotax. Sin. 9: 190. 1964; Crypsinus likiangensis (Ching) Nakaike; Phymatopteris likiangensis (Ching) Pichi Sermolli.
Rhizome ca. 4 mm in diam., densely scaly; scales brown, lanceolate, margin and both surfaces with fibrous hairs, apex acuminate. Fronds monomorphic. Stipe light brown, ca. 15 cm, glabrous; lamina pinnate near base, upper part pinnatisect, 2025 Χ 1520 cm, base cordate, margin entire, Lateral pinnae/lobes usually 3 or 4 pairs, ovate-lanceolate, 1012 Χ 23 cm, base contracted, widest at middle, apex acuminate. Costa and lateral veins distinct, veinlets obscure. Lamina papery, both surfaces glabrous. Sori round, in one row or two irregular rows on either side of costa, slightly closer to costa.
* Epiphytic on rocks in forests; ca. 2400 m. Yunnan (Lijiang).
三指假瘤蕨 san zhi jia liu jue
Polypodium trilobum Houttuyn, Nat. Hist. 14: 148. 1783; Crypsinus trilobus (Houttuyn) Copeland; Phymatodes triloba (Houttuyn) Ching; P. triphylla (Jacquin) C. Christensen & Tardieu; Phymatopsis triloba (Houttuyn) Ching; Phymatopteris triloba (Houttuyn) Pichi Sermolli; Polypodium incurvatum Blume; P. triphyllum Jacquin.
Check where this syn. came from
Rhizome 34 mm in diam., densely scaly throughout; scales castaneous at center, paler brown toward margin, ovate-lanceolate, rounded at peltate base, margin entire, apex acuminate. Fronds dimorphic, rarely intermediate with upper part fertile, lower part sterile. Sterile fronds: stipe light brown, 1020 cm, glabrous; lamina trilobed or pinnatifid, subdeltoid, rarely simple, 1020 Χ 1015 cm; margin entire, lateral lobes 13 pairs, ascending, broadly lanceolate, apical lobe broader, up to 45 cm wide; 1215 Χ 23 cm, apex acute or obtuse; costa raised, lateral veins distinct, veinlets hardly visible. Lamina leathery, both surfaces glabrous. Fertile fronds: stipe 2030 cm; lamina trilobed or pinnatisect with 24 pairs of lateral lobes; rachis narrowly winged or wingless in lower parts; lobes contracted, linear, less than 1 cm wide, apex acuminate. Sori immersed in deep cavities on abaxial side, raised on adaxial surface.
Epiphytic on tree trunks or on rocks; below 1300 m. Hainan [Indonesia, Malaysia, Philippines, Thailand, Vietnam].
十字假瘤蕨 shi zi jia liu jue
Polypodium cruciforme Ching, Sinensia 1: 47. 1930; Crypsinus cruciformis (Ching) Tagawa; Phymatodes cruciformis (Ching) Ching; Phymatopsis cruciformis (Ching) Ching; Phymatopteris cruciformis (Ching) Pichi Sermolli.
Rhizome 23 mm in diam., densely scaly throughout; scales light brown or glaucous, lanceolate, ca. 4 Χ 1 mm, margin entire or subentire, apex acuminate, long subulate. Fronds monomorphic. Stipe straw-colored, 25 cm, scaly at base, glabrous upward; lamina 3 or 5(7)-lobed, ca. 10 Χ 7.5 cm, base cuneate, margin minutely notched between veins, apex obtuse; basal lobes spreading, other lateral lobes slightly smaller and ascending, subdeltoid, gradually narrowed from base to acute apex; terminal lobe large, up to 8 Χ 2 cm. apex rounded. Costa and lateral veins raised on both surfaces, veinlets hardly visible. Lamina papery, both surfaces glabrous, abaxial surface glaucous. Sori round, in single row distal on lobes, slightly nearer to margin, sunken on abaxial surface and raised on adaxial surface.
Epiphytic on tree trunks in evergreen broad-leaved forests on hill tops. Guangdong (Longtoushan) [Thailand, N Vietnam].
昆明假瘤蕨 kun ming jia liu jue
Phymatopsis hirtella Ching, Acta Phytotax. Sin. 9: 191. 1964; Crypsinus hirtella (Ching) X. Cheng; Phymatopteris hirtella (Ching) Pichi Sermolli.
Rhizome 34 mm in diam., densely scaly; scales brown at central part, paler toward edges, ovate-lanceolate, ca. 4 Χ 11.5 mm, margin and abaxial surface whitish ciliate. Fronds monomorphic. Stipe straw-colored, 810 cm, sparsely hairy; lamina pinnatisect, 2025 Χ 1520 cm, base cordate. margin undulate and notched. Lateral lobes 46 pairs, ascending, lanceolate, 68 Χ 12 cm, apex acuminate. Costa and lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces pubescent. Sori round, large, medial.
* Terrestrial in Pinus yunnanensis forests; 20002100 m. Yunnan (Kunming, Yuanmou).
镰羽假瘤蕨 lian yu jia liu jue
Polypodium taeniatum Swartz, J. Bot. (Schrader) 1800[2]: 26. 1801; Crypsinus taeniatus (Swartz) Copeland; C. taeniatus var. palmatus C. Christensen [??not (Blume) Tagawa 1954]; Phymatodes falcatopinnata (Hayata) Ching; Phymatopsis falcatopinnata (Hayata) H. Itτ; P. taeniata (Swartz) Ching; P. taeniata var. palmata (Blume) Ching; Phymatopteris falcatopinnata (Hayata) S. G. Lu; P. palmata (Blume) Pichi Sermolli; P. taeniata (Swartz) Pichi Sermolli; Polypodium falcatopinnatum Hayata; P. palmatum Blume; S. falcatopinnata (Hayata) H. Ohashi & K. Ohashi; S. taeniata var. palmata (Blume) Parris.
Rhizome ca. 5 mm in diam., densely scaly; scales reddish brown or turning darker when old, lanceolate, broad at peltate base, margin entire or remotely denticulate, apex acuminate and hair-tipped. Fronds monomorphic. Stipe straw-colored, 1520 cm; lamina pinnately parted or pinnately divided, 2550 Χ 2030 cm, base cordate. margin notched or subentire. Lateral lobes/pinnae 2 or 3 pairs, opposite, linear-lanceolate, 1215 Χ 11.5 cm, ascending at base, falcate (bending downward) at upper portion, apex acuminate. Costa and lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, large, slightly closer to costa.
Taiwan (Lanyu Island) [Brunei, Indonesia, Malaysia, Philippines].
金平假瘤蕨 jin ping jia liu jue
Phymatopsis kingpingensis Ching, Acta Phytotax. Sin. 9: 191. 1964; Phymatopteris kingpingensis (Ching) Pichi Sermolli.
Rhizome 34 mm diam., with whitish bloom, sparsely scaly; scales black, lanceolate, margin whitish ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored to light brown, 1015 cm, glabrous; lamina pinnatisect, 1030 Χ 1520 cm, base rounded. margin sparsely notched. Lateral lobes 18 pairs, lanceolate, 810 Χ 11.5 cm, base usually contracted, apex acuminate. Lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, slightly nearer to costa.
* Epiphytic on tree trunks in evergreen broad-leaved forests; 20002100 m. Yunnan (Jingping).
耿马假瘤蕨 geng ma jia liu jue
Phymatodes connexa Ching, Bull. Fan Mem. Inst. Biol., n.s., 1: 306. 1949; Crypsinus connexus (Ching) X. Cheng; Phymatopsis connexa (Ching) Ching; Phymatopteris connexa (Ching) Pichi Sermolli; Pichisermollia connexa (Ching) Fraser-Jenkins; Pichisermollodes connexa (Ching) Fraser-Jenkins.
Rhizome ca. 3 mm in diam., densely scaly; scales yellowish brown, lanceolate, ca. 5 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored, 510 cm, glabrous; lamina pinnatisect, 1520 Χ 1015 cm, base broadly cuneate, margin notched, Lateral lobes usually 24 pairs, ascending, lanceolate, 58 cm Χ 812 mm, apex acuminate. Costa distinct, lateral veins and veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, medial.
* Epiphytic on tree trunks in forests; 25003000 m. Yunnan.
大围山假瘤蕨 da wei shan jia liu jue
Phymatopteris daweishanensis S. G. Lu, Fl. Reipubl. Popularis Sin. 6(2): 348. 2000; Crypsinus daweishanensis (S. G. Lu) X. Cheng.
Rhizome 34 mm in diam., densely scaly; scales whitish, lanceolate, margin with long whitish cilia, apex acuminate. Fronds close or remote, monomorphic. Stipe dark straw-colored, 810 cm, glabrous; lamina pinnatisect, ovate, 1517 Χ 1517 cm, base broadly cuneate to truncate, margin notched. Lateral lobes 3 or 4 pairs, lanceolate, 1015 Χ 12 cm, base slightly contracted, apex acute or obtuse. Costa and lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces glabrous, pale green. Sori round, large, very near margins.
* Epiphytic on tree branches in evergreen broad-leaved forests; ca. 1600 m. Yunnan (Daweishan, Pingbian).
黑鳞假瘤蕨 hei lin jia liu jue
Rhizome 36 mm in diam., very densely scaly; scales black at center, margins castaneous or dark brown, ovate-lanceolate, ca. 5 mm, stiff, margin ciliate, apex acute. Fronds monomorphic. Stipe straw-colored to light purplish, often inserted on pseudopodium 515 cm, densely scaly at base, more sparsely scaly upward; lamina pinnatisect to within 15 mm of costa, ovate in outline 2030(50) Χ 825 cm, base cordate. margin notched, Lateral lobes usually 310 pairs, lower 1 or 2 pairs deflexed, others spreading or ascending, lanceolate, 515 Χ 12 cm, apex acuminate, Costa and lateral veins distinct, veinlets obscure. Lamina herbaceous, abaxial surface pale green and sparsely scaly, scales brown or light brown, ovate, thin, mainly along costa, adaxial surface green, glabrous or papillate-pubescent on costa and main veins. Sori round, slightly near costa.
Terrestrial or epiphytic on tree trunks or on rocks; 19003500 m. Hunan, Sichuan, Xizang, Yunnan [Bhutan, NE India, Nepal, Thailand].
1a. Rachis and costae glabrous ............................................................................................................. 30a. var. ebenipes
1b. Rachis and costae pubescent ............................................................................................................ 30b. var. oakesii
黑鳞假瘤蕨 (原变种) hei lin jia liu jue (yuan bian zhong)
Polypodium ebenipes Hooker, Sp. Fil. 5: 88. 1864; Crypsinus ebenipes (Hooker) Copeland; C. ebenipes var. subebenipes (Ching) K. Iwatsuki et al.; C. nepalensis (Nakaike) Subh. Chandra [??okay]; Phymatodes ebenipes (Hooker) Ching; Phymatopsis ebenipes (Hooker) J. Smith; P. ebenipes var. subebenipes (Ching) K. Iwatsuki[??not in Tropicos]; P. subebenipes Ching; Phymatopteris ebenipes (Hooker) Pichi Sermolli; P. nepalensis (Nakaike) Subh. Chandra; P. subebenipes (Ching) Pichi Sermolli; Pichisermollia ebenipes (Hooker) Fraser-Jenkins; P. subebenipes (Ching) Fraser-Jenkins; Pichisermollodes ebenipes (Hooker) Fraser-Jenkins; P. subebenipes (Ching) Fraser-Jenkins; Pleopeltis ebenipes (Hooker) Beddome.
Rachis and costae glabrous.
Terrestrial or epiphytic on tree trunks or on rocks; 19003200 m. Hunan, Sichuan, Xizang, Yunnan [Bhutan, NE India, Nepal, Thailand].
毛轴黑鳞假瘤蕨 mao zhou hei lin jia liu jue
Polypodium ebenipes var. oakesii C. B. Clarke, Trans. Linn. Soc. London, Bot. 1: 564. 1880; Phymatopteris ebenipes var. oakesii (C. B. Clarke) Satija & Bir.
Rachis and costa pubescent.
Epiphytic on rocks; 23003500 m. Xizang, Yunnan [NE India].
大叶玉山假瘤蕨 da ye yu shan jia liu jue
Phymatodes echinospora Tagawa, Acta Phytotax. Geobot. 3: 95. 1934; Crypsinus echinosporus (Tagawa) Tagawa; Phymatopsis echinospora (Tagawa) H. Itτ; Phymatopteris echinospora (Tagawa) Pichi Sermolli.
Rhizome 812 mm in diam., densely scaly throughout; scales dark brown at center, margins brown, lanceolate, 67 Χ ca. 1 mm, rounded at base, margin entire, apex acuminate. Fronds monomorphic. Stipe straw-colored, 515 cm, glabrous; lamina pinnatisect, 1520 Χ 1015 cm, base truncate or cordate. margins notched. Lateral lobes 511 pairs, lowest pair deflexed, upper pairs spreading, linear-lanceolate, 510 cm Χ 818 mm, apex acuminate. Costa and lateral veins distinct, raised on both surfaces, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, large, close to edges, sunken in cavities on abaxial surface and raised on adaxial surface.
* Epiphytic on tree trunks or on rocks; 20002400 m. Taiwan.
展羽假瘤蕨 zhan yu jia liu jue
Polypodium quasidivaricatum Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30: 446. 1911; Crypsinus intermedius (Ching) Tagawa; C. quasidivaricatus (Hayata) Copeland; Phymatodes intermedia Ching; P. quasidivaricata (Hayata) H. Itτ [??Ching]; Phymatopsis intermedia (Ching) Ching; P. quasidivaricata (Hayata) H. Itτ; Phymatopteris intermedia (Ching) Pichi Sermolli; P. quasidivaricata (Hayata) Pichi Sermolli; Pichisermollia quasidivaricata (Hayata) Fraser-Jenkins; Pichisermollodes quasidivaricata (Hayata) Fraser-Jenkins; Polypodium divaricatum Hayata (Bot. Mag. (Tokyo) 23: 78. 1909), not E. Fournier (1872); P. morianum C. Christensen.
Rhizome ca. 3 mm in diam., densely scaly throughout; scales castaneous at center, margins brown or light brown, linear-lanceolate, ca. 5 Χ 1 mm, margin long ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 510 cm, glabrous; lamina pinnatisect, 1020 Χ 515 cm, base cordate, margin notched or serrulate. Lateral lobes 25 pairs, lowest pair usually much deflexed, upper pairs spreading or ascending, linear-lanceolate, 57 Χ 11.5 cm, base slightly contracted, apex acuminate. Costa and lateral veins distinct on both surfaces, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, slightly near costa, superficial.
* Epiphytic on tree trunks or on rocks in forests; ca. 1000 m. Taiwan.
Selliguea quasidivaricata is very close to S. laciniata (Beddome) Hovenkamp, which is widely distributed from Thailand to New Guinea.
The name Phymatopteris quasidivaricata has been applied erroneously to a number of specimens from high mountains in Yunnan and possibly other provinces on the mainland. Shieh & [??] (in H. L. Li et al., Fl. Taiwan 1: 177. 1975) misidentified material of this species as Crypsinus veitchii (Baker) Copeland.
Orthography: albipes??
灰鳞假瘤蕨 hui lin jia liu jue
Polypodium albopes C. Christensen & Ching, Bull. Dept. Biol. Sun Yatsen Univ. 6: 15. 1933; Crypsinus albopes (C. Christensen & Ching) Tagawa; C. chinensis (Ching) Tagawa; Phymatodes albopes (C. Christensen & Ching) Ching; P, chinensis Ching; Phymatopsis albopes (C. Christensen & Ching) Ching; Phymatopteris albopes (C. Christensen & Ching) Pichi Sermolli; Pichisermollodes albopes (C. Christensen & Ching) Fraser-Jenkins.
Rhizome 45 mm in diam., densely scaly throughout; scales whitish or light brown, dark when older, linear-lanceolate, 67 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe light brown, 812 cm, glabrous; lamina pinnatisect, 2025 Χ 1015 cm, base truncate or cordate, margin notched. Lateral lobes 810 pairs, lowest pair deflexed, upper pairs spreading or ascending, lanceolate, 58 Χ 11.5 cm, apex acuminate. Costa distinct and raised on both surfaces, lateral veins and veinlets obscure. Lamina papery, both surfaces glabrous. Sori round, near or slightly near edges, sunken on abaxial surface and raised on adaxial surface.
* Epiphytic on tree trunks. Fujian, Guangdong, Guangxi (Yaoshan), Hunan, Jiangxi, Yunnan.
片马假瘤蕨 pian ma jia liu jue
Phymatopteris pianmaensis W. M. Chu, Acta Bot. Yunnan., Suppl. 5: 56. 1992; Crypsinus pianmaensis (W. M. Chu) X. Cheng.
Rhizome 45 mm in diam., densely scaly throughout; scales brown at peltate center, whitish on other parts, lanceolate, ca. 10 Χ 2 mm, margin entire, apex acuminate. Fronds monomorphic. Stipe straw colored, 510 cm, glabrous; lamina pinnatisect, or partly pinnate with free lower lobes, 1020 Χ 512 cm, base truncate or cordate, margin sparsely notched. Lateral lobes 58 pairs, lowest pair deflexed, upper pairs spreading, slightly contracted at base, lanceolate, 26 Χ 11.8 cm, apex obtuse. Costa distinct, lateral veins slender, hardly visible, veinlets obscure. Lamina herbaceous, both surfaces pubescent. Sori round, slightly near costa.
* Epiphytic on tree trunks or on rocks; ca. 2100 m. Yunnan (Lushui, Pingbian).
毛叶假瘤蕨 mao ye jia liu jue
Polypodium shensiense H. Christ var. nigrovenium H. Christ, Bull. Acad. Int. Gιogr. Bot. 15: 106. 1906; Crypsinus nigrovenia (H. Christ) K. Iwatsuki; Phymatodes nigrovenia (H. Christ) Ching; Phymatopsis laipoensis Ching; P. nigrovenia (H. Christ) Ching; P. veitchii (Baker) H. Itτ var. nigrovenium (H. Christ) Takeda[??not in Tropicos]; Phymatopteris laipoensis (Ching) Pichi Sermolli; P. nigrovenia (H. Christ) Pichi Sermolli; Pichisermollia nigrovenia (H. Christ) Fraser-Jenkins; Pichisermollodes nigrovenia (H. Christ) Fraser-Jenkins; Polypodium nigrovenium (H. Christ) Ching (1930), not H. Christ (1896).
Rhizome ca. 2 mm in diam., with whitish bloom, sparsely scaly; scales black or dark brown on young parts, lanceolate, ca. 3 mm, margin sparsely ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 46 cm, slender, glabrous; lamina pinnatisect, 810 Χ 56 cm, base shallowly cordate. margin shallowly serrate, Lateral lobes 35 pairs, lowest pair deflexed, lanceolate, 34 Χ ca. 1 cm, base usually contracted, apex obtuse or acute. Veins distinct on both surfaces, lateral veins distinct, tortuous. Lamina papery, abaxial surface glabrous, adaxial surface pubescent. Sori round, near costa.
* Epiphytic on tree trunks or on rocks; 25003300 m. Hubei, Sichuan (Emei Shan), Yunnan.
圆齿假瘤蕨 yuan chi jia liu jue
Phymatopteris incisocrenata Ching ex W. M. Chu & S. G. Lu, Fl. Reipubl. Popularis Sin. 6(2): 348. 2000; Crypsinus incisocrenatus (Ching ex W. M. Chu & S. G. Lu) X. Cheng.
Rhizome ca. 2 mm in diam., with whitish bloom, scaly; scales dark brown, lanceolate, ca. 4 mm, margin ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 1015 cm, glabrous; lamina pinnately parted[????], 1215 Χ 1012 cm, base shallowly cordate, margin crenate-serrate. Lateral lobes 47 pairs, lowest pair deflexed, upper pairs spreading or ascending, lanceolate, 46 Χ 1.52 cm, apex acute. Veins distinct on both surfaces, lateral veins distinct[????], tortuous. Lamina papery, abaxial surface glabrous, adaxial surface pubescent. Sori round, near costa.
* Epiphytic on tree trunks or on rocks; 25003100 m. Yunnan.
陕西假瘤蕨 shan xi jia liu jue
Polypodium shensiense H. Christ, Nuovo Giorn. Bot. Ital., n.s., 4: 99. 1897; Crypsinus shensiensis (H. Christ) X. Cheng; Phymatopsis shensiensis (H. Christ) Ching; Phymatopteris shensiensis (H. Christ) Pichi Sermolli; Polypodium senanense Maximowicz; P. shensiense var. filipes H. Christ.
Rhizome 1.52 mm in diam., densely scaly throughout; scales brown or dark brown at peltate base, ovate-lanceolate, margin sparsely ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored or dark stramineous, 25 cm, slender, glabrous; lamina pinnatisect, 510 Χ 57 cm, base truncate or shallowly cordate. margin shallowly serrate. Lateral lobes 25 pairs, lowest pair slightly deflexed at base, 23 Χ ca. 1 cm, base usually contracted, apices obtuse or acute. Costa and lateral veins distinct on both surfaces, veinlets hardly visible. Lamina herbaceous, both surfaces glabrous, grayish green, Sori round, slightly near costa.
· Epiphytic on tree trunks or on rocks, rarely terrestrial; 13003600 m. Henan, Shaanxi, Shanxi, Sichuan, Xizang, Yunnan.
Crypsinus veitchii (Baker) Copeland from Japan and Korea is closely related to Selliguea shensiense but the lamina lacks the deflexed basal lobes [??citation {Iwatsuki, Fl.Japan}].
紫柄假瘤蕨 zi bing jia liu jue
Polypodium crenatopinnatum C. B. Clarke, J. Linn. Soc., Bot. 25: 99. 1889; Phymatodes crenatopinnata (C. B. Clarke) Ching; Phymatopsis crenatopinnata (C. B. Clarke) Ching; Phymatopteris crenatopinnata (C. B. Clarke) Pichi Sermolli; P. pseudoserratum H. Christ[??not in IPNI or Tropicos]; Pichisermollia crenatopinnata (C. B. Clarke) Fraser-Jenkins; Pichisermollodes crenatopinnata (C. B. Clarke) Fraser-Jenkins; Pleopeltis crenatopinnata (C. B. Clarke) Beddome, [crenato-pinnata]; Polypodium connatum H. Christ; P. pseudoserratum H. Christ.
Rhizome ca. 2 mm in diam., densely scaly throughout; scales black at center, margins brown, lanceolate, 23 mm, margin ciliate, apex acuminate. Fronds monomorphic. Stipe purplish or castaneous, 1020 cm, glabrous; lamina pinnately (to partially bipinnately) divided almost to midrib, deltoid-ovate in outline, 520 Χ 510 cm, base truncate. Lateral lobes 38 pairs, far apart, 510 Χ 0.51.2 cm, base distinctly contracted, decurrent into very narrowly winged rachis, margin very shallowly crenate to irregularly incised or lobed, sometimes quite deeply lobed in more basal lobes, apex acute or obtuse. Veins distinct, veinlets usually obscure. Lamina papery, both surfaces glabrous. Sori round or elliptic, medial or slightly nearer to costa.
Terrestrial in forests; 19002900 m. Guangxi, Guizhou, Hunan, Sichuan, Xizang, Yunnan [NE India].
刺齿假瘤蕨 ci chi jia liu jue
Polypodium glaucopsis Franchet, Bull. Soc. Bot. France 32: 29. 1885; Crypsinus glaucopsis (Franchet) Tagawa; Phymatodes veitchii (Baker) Ching var. glaucopsis (Franchet) Ching; Phymatopsis glaucopsis (Franchet) Ching; Phymatopteris glaucopsis (Franchet) Pichi Sermolli; Pleopeltis glaucopsis (Franchet) Beddome; Polypodium veitchii Baker var. glaucopsis (Franchet) C. Christensen ex Handel-Mazzetti.
Rhizome ca. 2 mm in diam., densely scaly throughout; scales castaneous at center, margins brown, lanceolate, 23 mm, margin ciliate, apex acuminate. Fronds monomorphic. Stipe brown, 515 cm, glabrous; lamina pinnatisect, 715 Χ 510 cm, base cordate, margin sharply serrulate. Lateral lobes 2 or 3 pairs, closely spaced, 35 Χ 12 cm, contracted at base, widest at middle, apex obtuse. Lateral veins tortuous, not ending at margin, veinlets distinct[????]. Lamina papery, both surfaces glabrous. Sori round, slightly near costa.
Terrestrial or epiphytic on rocks; 27003700 m. Sichuan, Yunnan [N India].
钝羽假瘤蕨dun yu jia liu jue
Phymatodes conmixta Ching, Bull. Fan Mem. Inst. Biol., n.s. 1: 307. 1949; Crypsinus conmixtus (Ching) X. Cheng; Phymatopsis conmixta (Ching) Ching; Phymatopteris conmixta (Ching) Pichi Sermolli.
Rhizome ca. 2 mm in diam., densely scaly throughout; scales light brown on young parts, dark brown when older, lanceolate, margin ciliate, apex acuminate. Fronds monomorphic. Stipe purplish, 510 cm, glabrous; lamina partly pinnate, upper part pinnatisect, 1015 Χ 58 cm, base shallowly cordate. margin densely serrulate or duplicate-serrulate. Lateral lobes 57 pairs, far apart, linear, 35 Χ 0.51 cm, usually not contracted at base, apex obtuse. Costa and lateral veins distinct, veinlets hardly visible. Lamina herbaceous, both surfaces glabrous. Sori elliptic or round, medial or slightly near costa.
· Terrestrial or epiphytic on rocks in forests; 31003600 m. Sichuan, Yunnan.
斜下假瘤蕨 xie xia jia liu jue
Phymatodes stracheyi Ching, Contr. Inst. Bot. Natl. Acad. Peiping 2(3): 83. 1933; Crypsinus stracheyi (Ching) Panigrahi & Patnaik; Phymatopsis stracheyi (Ching) Ching; Phymatopteris stracheyi (Ching) Pichi Sermolli; Polypodium stracheyi (Ching) C. Christensen[??okay].
In or ex C. Christensen?
Rhizome ca. 3 mm in diam., densely scaly throughout; scales castaneous at center, margin and apex brown, lanceolate, margin ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 58 cm, glabrous; lamina pinnatisect, 1012 Χ 1012 cm, base cordate. margin densely serrulate. Lateral lobes 24 pairs, lowest pair deflexed, lanceolate, 57 Χ 11.5 cm, widest at base, narrowed from base, apex acuminate. Lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, near costa.
Epiphytic on tree trunks; 28003700 m. Hubei, Sichuan, Xizang, Yunnan [Bhutan, N India, Nepal].
Baker (J. Bot. 1889: 177. 1889) treated material of this species (Selliguea stracheyi) as Polypodium malacodon Hooker; C. B. Clarke (Trans. Linn. Soc. London, Bot. 1: 563. 1880) included material of this species within Polypodium stewartii C. B. Clarke.
Polypodium trifidum H. Christ (1905) not Don (1825), Franchet & Savatier (1876), etc.
交连假瘤蕨 jiao lian jia liu jue
Phymatopsis conjuncta Ching, Acta Phytotax. Sin. 9: 196. 1964; P. wuyishanica Ching & K. H. Shing; Phymatopteris conjuncta (Ching) Pichi Sermolli.
Rhizome ca. 3 mm in diam., densely scaly throughout; scales usually black at center, margins brown or pale brown, lanceolate, 45 mm, margin ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 510 cm, glabrous; lamina pinnatisect, 1015 Χ 612 cm, base cordate, margin sharply serrate. Lateral lobes 24 pairs, lowest pair deflexed, ovate-lanceolate, 58 Χ 1.52 cm, base slightly contracted, usually broadest at middle, apex acute or obtuse. Lateral veins distinct, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, near costa.
· Epiphytic on tree trunks or on rocks; 15003600 m. Anhui, Fujian, Guangxi, Guizhou, Henan, Hubei, Hunan, Shaanxi, Sichuan, Xizang, Yunnan.
尾尖假瘤蕨 wei jian jia liu jue
Pleopeltis stewartii Beddome, Ferns Brit. India 2: 204. 1866[??1867]; Crypsinus stewartii (Beddome) Copeland; Phymatodes stewartii (Beddome) Ching; Phymatopsis stewartii (Beddome) Ching; Phymatopteris stewartii (Beddome) Pichi Sermolli; Pichisermollia stewartii (Beddome) Fraser-Jenkins; Pichisermollodes stewartii (Beddome) Fraser-Jenkins; ?Polypodium cyrtolobum J. Smith ex C. B. Clarke; ?P. malacodon Hooker var. majus J. Smith ex Hooker; P. stewartii (Beddome) Baker[??okay].
Rhizome 34 mm in diam., with whitish bloom, scaly; scales uniformly castaneous or black, lanceolate, acumen whitish ciliate. Fronds remote or close. Stipe light brown, 710 cm, glabrous; lamina pinnatisect, 1530 Χ 1018 cm, base rounded or shallowly cordate, margin regularly obtusely serrate, rarely basal lobe forked. Lateral lobes 24 pairs, 1015 Χ 12 cm, spreading and slightly contracted at base, from middle part curved toward apex of lamina, apex caudate-acuminate. Lateral veins distinct, not reaching margin, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, near costa.
Epiphytic on tree trunks or on rocks; 24003000 m. Sichuan, Xizang, Yunnan [N India, Nepal].
乌鳞假瘤蕨 wu lin jia liu jue
Phymatopsis nigropaleacea Ching, Acta Phytotax. Sin. 9: 196. 1964; Crypsinus nigropaleaceus (Ching) Nakaike; Phymatopteris nigropaleacea (Ching) S. G. Lu; P. stewartii[??] var. nigropaleacea (Ching) X. Cheng[??not in Tropicos].
Rhizome 34 mm diam., with whitish bloom, scaly; scales black, lanceolate, 45 mm, margin densely whitish ciliate, apex acuminate. Fronds close or remote. Stipe light purplish, 510 cm, glabrous; lamina pinnatisect, or partly pinnate with free lower lobes, 1535 Χ 715 cm, base rounded, margin stramineous, sharply duplicate-serrate, 35-toothed between adjacent lateral veins. Lateral lobes 49 pairs, lanceolate, 58 Χ 11.5 cm, curved toward apex of lamina, apex acuminate. Lateral veins distinct, ending before margin, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, slightly near costa.
* Epiphytic on tree trunks or on rocks; 26003800 m. Sichuan, Yunnan.
紫边假瘤蕨 zi bian jia liu jue
Phymatodes roseomarginata Ching, Bull. Fan Mem. Inst. Biol., ser. 2, 1: 305. 1949; Phymatopsis roseomarginata (Ching) Ching; Phymatopteris roseomarginata (Ching) Pichi Sermolli.
Rhizome ca. 3 mm in diam., with whitish bloom, scaly; scales black, lanceolate, margin whitish ciliate, apex acuminate. Fronds remote or close. Stipe light brown, 24 cm, slender, glabrous; lamina pinnatisect, 510 Χ 57 cm, base rounded or broadly cuneate, margin densely sharply serrulate, cartilaginous, usually purple or reddish. Lateral lobes 2 or 3 pairs, curving toward apex of lamina, 24 Χ 0.51 cm, base decurrent, apex acuminate or caudate. Lateral veins distinct, ending before margin, veinlets obscure. Lamina herbaceous, both surfaces glabrous. Sori round, medial.
* Epiphytic on tree trunks or on rocks; ca. 3200 m. Yunnan.
西藏假瘤蕨 xi zang jia liu jue
Phymatopsis tibetana Ching & S. K. Wu, Fl. Xizang. 1: 325. 1983; Crypsinus tibetanus (Ching & S. K. Wu) Nakaike; Phymatopteris tibetana (Ching & S. K. Wu) W. M. Chu; Pichisermollia tibetana (Ching & S. K. Wu) Fraser-Jenkins; Pichisermollodes tibetana (Ching & S. K. Wu) Fraser-Jenkins.
Rhizome 34 mm in diam., densely scaly throughout; scales black at peltate center, reddish brown on other parts, ovate-lanceolate, margin ciliate, apex acuminate. Fronds monomorphic. Stipe light purplish, 1520 cm, glabrous; lamina pinnatisect, oblong-deltoid, 1520 Χ 1518 cm, base truncate or shallowly cordate, margin shallowly serrate. Lateral lobes 36 pairs, lanceolate, 810 Χ 1.52 cm, apex acuminate. Lateral veins distinct on both surfaces and ending at margin, veinlets hardly visible. Lamina herbaceous, both surfaces glabrous, abaxial surface pale green. Sori round, near costa.
* Epiphytic on tree trunks; 24003400 m. Xizang, Yunnan.
The type is from Xizang.
芒刺假瘤蕨 mang ci jia liu jue
Polypodium malacodon Hooker, Sp. Fil. 5: 87. 1864[??1863]; Crypsinus cartilagineoserratus (Ching & S. K. Wu) Nakaike; C. malacodon (Hooker) Copeland; Phymatodes malacodon (Hooker) Ching; Phymatopsis cartilagineoserrata Ching & S. K. Wu, [cartilagineo-serrata]; P. malacodon (Hooker) Ching; Phymatopteris cartilagineoserrata (Ching & S. K. Wu) S. G. Lu; P. malacodon (Hooker) Pichi Sermolli; Pichisermollia malacodon (Hooker) Fraser-Jenkins; Pichisermollodes malacodon (Hooker) Fraser-Jenkins; Pleopeltis malacodon (Hooker) Beddome (1883), not Beddome (1876).
Rhizome ca. 3 mm in diam., densely scaly throughout; scales black or dark brown at center, pale brown toward edges, lanceolate, ca. 4 mm, margin toothed, apex acuminate. Fronds monomorphic. Stipe purplish or straw-colored, 510 cm, glabrous; lamina pinnatisect, 1015 Χ 814 cm, base cordate. margin serrulate or duplicate-serrate, with sharply pointed teeth. Lateral lobes usually 13 pairs, lowest pair slightly deflexed at base and bending upward at apex, 57 Χ 1.52 cm, apex acute. Midrib and lateral veins usually tortuous, veinlets hardly visible. Lamina subleathery, both surfaces glabrous, abaxial surface pale green, adaxial surface green. Sori round, medial or slightly near costa.
* Epiphytic on rocks; ca. 3200 m. Xizang.
The type of Phymatopsis cartilagineoserrata is a good match with that of Selliguea malacodon which has rhizome scales with a light ciliate margin, and curved pinnae with sharply aristate teeth. Chinese material previously named as Phymatopteris malacodon is here included within the following species, Selliguea albidoglauca.
Hovenkamp: I have a number of specimens from Nepal and Bhutan of P. malacodon, and I find that in all cases, they have scales with a light ciliate margin, and curved pinnae with sharply aristate teeth. According to your key and description, this combination is not allowed in P. malacodon, but would indicate that they belong to P. cartilagineo-serrata. There can be little doubt that the sharply toothed specimens represent the true P. malacodon, and that P. cartilagineoserrata is therefore a synonym. I cannot assess the affinities of the specimens here considered as P. malacodon, but I think they may have to be called P. austrosinicum.
弯弓假瘤蕨 wan gong jia liu jue
Polypodium albidoglaucum C. Christensen, Index Filic., Suppl.: 58. 1912; P. austrosinicum H. Christ (Bull. Acad. Int. Gιogr. Bot. 16: 107. 1906), not H. Christ (in C. Christensen, 1906).
Rhizome ca. 3 mm in diam., densely scaly; scales dark brown at center, light brown at margin, shiny, ovate-lanceolate, margin ciliate, apex acuminate. Fronds monomorphic. Stipe straw-colored, 14 cm, slender, glabrous; lamina pinnatisect, triangular in outline, 48 Χ 36 cm, base cordate, margin with up to 2 mm aristate teeth, Lateral lobes 1 or 2 pairs, lowest pair curved, deflexed at base, bending at apex, spreading or slightly ascending further up, 23 Χ 11.5 cm, apex obtuse or acute. Lateral veins distinct, veinlets hardly visible. Lamina herbaceous, both surfaces glabrous. Sori near costa.
Epiphytic on tree trunks or on rocks; 28003700 m. Sichuan, Xizang, Yunnan [Bhutan, N India, Nepal].
The material here included within Selliguea albidoglauca was treated as Phymatopteris malacodon (= Selliguea malacodon) in FRPS. It differs from the type of that species by the lack of the aristate marginal teeth of the lamina characteristic of S. malacodon s. str. Chinese material of the true S. malacodon was named P. cartilagineoserrata.
The choice of epithet for this taxon depends on the priority of Polypodium austrosinicum H. Christ versus P. austrosinicum H. Christ ex C. Christensen (a replacement name for P. henryi H. Christ, here treated as a synonym of Microsorum fortunei (T. Moore) Ching), both published in 1906. The only direct indication of priority is the action of Christensen who claimed priority for the latter, published in his own work, and published the replacement name P. albidoglaucum for the former. In the absence of other information on the exact publication dates, this has to be accepted and the correct epithet for this taxon is that of Christensen.
修蕨xiu jue
Polypodium feei (Bory) Mettenius; P. pedunculatioides Ching.
Rhizome ca. 4 mm in diam., densely scaly when young; scales reddish brown, ovate-lanceolate, 68 mm, peltate at base, margin entire, apex acuminate. Fronds remote, slightly dimorphic. Sterile fronds: stipe 1820 cm. Lamina 1520 Χ 79 cm, base broadly cuneate, margin entire, apex acute. Veins reticulate, lateral veins raised on abaxial surface, veinlets obscure. Lamina leathery, both surfaces glabrous. Fertile fronds: stipe 2035 cm; lamina ovate, 1320 Χ ca. 3 cm, base cuneate, apex acuminate. Sori reddish brown, linear, ca. 3 mm in diam., borne between adjacent lateral veins.
Epiphytic on tree trunks; below 1200 m. Guangdong [Indonesia, Malaysia, Philippines to[??list countries] Polynesia].
锡金假瘤蕨属 xi jin jia liu jue shu
Rhizome long-creeping, 35 mm in diam. Fronds 13 cm apart; rhizome scales linear-lanceolate, central portion castaneous, margin brown and somewhat ciliolate, apex acuminate. Fronds simple; stipe straw-colored, 515 cm, very slender; lamina deeply pinnatifid, rachis narrowly winged or lower part wingless; lobes usually 15 Χ 0.81.5 cm, apex acute, with conspicuous main veins and distinct lateral veins, costal areoles in 1 or 2 series, lowest lateral lobe somewhat decurrent; venation type goniophlebioid; margins with distinct small incisions or notches; texture subleathery, with thick multicellular trichomes on both surfaces; apical part of lamina fertile, sori in a single series in lower areoles.
Monotypic: China, India (Sikkim), Nepal.
The extensive generic synonymy [??where] is a clear indication of the problems that there have been in placing this species within a genus.
锡金假瘤蕨 xi jin jia liu jue
Polypodium erythrocarpum Mettenius ex Kuhn, Linnaea 36: 135. 1869; Crypsinus erythrocarpus (Mettenius ex Kuhn) Tagawa; Goniophlebium erythrocarpum (Mettenius ex Kuhn) Beddome; Phymatodes erythrocarpa (Mettenius ex Kuhn) Ching; Phymatopsis erythrocarpa (Mettenius ex Kuhn) Ching; Phymatopteris erythrocarpa (Mettenius ex Kuhn) Pichi Sermolli; Pichisermollia erythrocarpa (Mettenius ex Kuhn) Fraser-Jenkins; Pichisermollodes erythrocarpa (Mettenius ex Kuhn) Fraser-Jenkins.
Rhizome 23 mm in diam., covered with whitish bloom and sparse scales; scales black or dark brown, lanceolate, ciliate at margin, acuminate at apex. Fronds monomorphic, deciduous. Stipe straw-colored, 510 cm, pubescent. Lamina pinnately parted[????] or pinnately divided in lower part, 1015 Χ 510 cm, cordate at base. Lateral lobes 57(10) pairs, lanceolate, 25 Χ ca. 1 cm, slightly contracted at base, crenate-serrate and ciliate at margin, obtuse or acute at apex. Costa and lateral veins distinct, veinlets obscure. Texture papery, green, densely hairy on both surfaces. Sori round, slightly near costa.
Epiphytic on tree trunks; 27002800 m. Xizang (Medog) [India (Sikkim), Nepal].
槲蕨属 hu jue shu
Polypodium subg. Drynaria Bory, Ann. Sci. Nat. 5: 463. 1825.
Zhang Xianchun (张宪春)1;
Plants epiphytic, epilithic, or terrestrial. Rhizome short to long-creeping; rhizome scales appressed or, often squarrosely, spreading, basifixed or peltate, margin toothed; fronds usually dimorphic, with base and foliage frond, rarely monomorphic (basal frond absent); basal fronds sessile, orbicular to ovate-elliptical, entire to lobed up to 2/3; foliage fronds internally monomorphic or apical fertile part slightly narrowed, stalked, pinnatifid, apex often aborted, with a lateral pinna taking its place, rarely pinnate (then pinnae articulate to rachis, deciduous), hairs sometimes present, spread throughout lamina; in pinnatifid species, pinnae abscissile[??spelling] from costa and from each other by a line of abscission between costa and base of sinus; basal pinnae reduced; costae, costules, and main veins prominent abaxially, cross-veins and minor venation evident, much anastomosing with occasional free-included veinlets, veins forming glandular plexuses[??spelling] along costa, usually just above costules; mature fronds without hydathodes on adaxial surface, margins cartilaginous, lamina of mature fronds scaly at first, glabrescent. Sori small, in rows along veins or connecting veins, very shallowly impressed. Sporangia glabrous or sometimes with glandular hairs. Spores with spines or small globules. Chromosome number n = 36, 37. [??x]
Sixteen species: paleotropical, mainly in subtropical and tropical regions, epiphytic, epilithic, or terrestrial; nine species (one endemic) in China, one (D. sinica) extends to N China [??North of China].
1a...... Foliage fronds pinnate; pinnae articulate to rachis ............................................................................ 1. D. rigidula
1b..... Foliage fronds pinnatifid; pinnae not articulate to rachis.
2a...... Basal fronds thick, not transparent; sori in two or more rows between costa and margin.
3a...... Basal fronds rounded, margin entire or shallowly undulate; sori small, irregularly spaced between lateral veins 2. D. bonii
3b..... Basal fronds ovate or cordate, margin distinctly lobed; sori large, in 1 or 2 rows between lateral veins.
4a. Plants 3040 cm high; basal fronds 35 Χ 24 cm; sori in one row between lateral veins .. 3. D. roosii
4b..... Plants up to 100 cm high; basal fronds 2040 Χ 1632 cm; sori in two rows between lateral veins 4. D. quercifolia
2b..... Basal fronds thin, transparent, sometimes absent (D. parishii); sori in one row between costa and margin.
5a...... Scales peltate, rigid and imbricate; foliage fronds glabrous.
6a. Basal fronds absent; foliage fronds with 59 pairs of pinnae .............................................. 5. D. parishii
6b..... Basal fronds present; foliage fronds with 812 pairs of pinnae ..................................... 6. D. propinqua
5b..... Scales basifixed, soft and tufted; foliage fronds often hairy.
7a. Apex of foliage fronds not aborted; margin of lamina entire, long ciliate .............................. 7. D. mollis
7b..... Apex of foliage fronds aborted; margin of lamina serrate or sinuate, glabrous or with short hairs in sinus.
8a...... Basal fronds oblong-lanceolate, 515 Χ 36 cm; foliage fronds 712 cm wide; pinnae 0.51.2 cm wide, apex of lower pinnae rounded, of upper pinnae acute ............................................................................... 8. D. baronii
8b..... Basal fronds ovate, 614 Χ 49 cm; foliage fronds 1220 cm wide; pinnae 1.22 cm wide, apex of pinnae acuminate ................................................................................................................................................... 9. D. delavayi
硬叶槲蕨 ying ye hu jue
Polypodium rigidulum Swartz, J. Bot. (Schrader) 1800(2): 26. 1801; Drynaria baudouinii E. Fournier; D. diversifolia (R. Brown) J. Smith; D. gaudichaudii (Bory) Gaudichaud; D. pinnata Fιe; D. rigidula var. koordersii Alderwerelt; Goniophlebium rigidulum (Swartz) T. Moore; Phymatodes gaudichaudii (Bory) C. Presl; Polypodium baudouini (E. Fournier) Baker; P. diversifolium R. Brown (1810), not Willdenow (1810); P. gaudichaudii Bory; P. rigidulum var. vidgeni F. M. Bailey; P. speciosum Blume.
Rhizome shortly creeping, 12 cm in diam.; scales brown to dark brown, with a lighter margin, spreading, peltate, 513 Χ 0.51.5 mm, margin ciliate, apex acute to acuminate; fronds dimorphic, rachises persistent; basal fronds 1030 Χ 515 cm, lobed from 1/34/5, margin irregularly and finely denticulate; foliage fronds stalked, stipe up to 40 cm, not winged, with two rows of nectaries; lamina pinnate, 25100(200) Χ 1250 cm, apex aborted; pinnae articulate to rachis, all equally long or smaller toward apex, 825(30) Χ 0.51.5 cm, base narrowly cuneate, margin crenate to serrate, apex obtuse to acuminate, free veinlets simple or absent, hydathodes absent. Sori in one row between costa and margin, costal, single between veins, round, sunken. Spores with short, blunt spines and globules.
Epiphytic, encircling tree trunks many times, rarely epilithic; sea level up to 2000(2400) m. Hainan, SW Yunnan [Indochina [??countries], Myanmar, Thailand; tropical Australia, Malesia, Polynesia].
Drynaria rigidula differs from all other species of Drynaria by the pinnate fronds. It is the sole member of Sect. Poronema, to appropriately show its isolated position.
团叶槲蕨 tuan ye hu jue
Drynaria meeboldii Rosenstock.
Rhizome shortly creeping, terete to dorsiventrally flattened, ca. 3 mm in diam., 12 cm wide[??okay]; scales squarrose or spreading from an appressed base, peltate, 212 Χ 1.53 mm, abruptly contracted from a broad base, often with a distinct, brown midrib, margin irregularly and shortly dentate at base, acumen often subulate and nearly entire; fronds dimorphic, glabrous; basal fronds contiguous or overlapping, often completely covering and surrounding rhizome, sessile, (4)1015 Χ (3.5)812 cm, nearly entire to shallowly lobed; foliage fronds stalked, stipe up to 20 cm, conspicuously winged with a sinuose wing; lamina pinnatifid up to ca. 1 mm from costa, 3070 Χ 2030 cm, apex aborted; pinnae 37 pairs, slightly ascending, 1.53 cm distant, broadly lanceolate, 720 Χ 2.55 cm, base slightly narrowed and decurrent, margin subentire to shallowly sinuate, apex obtuse to long acuminate. Sori in 2 irregular rows between costa and margin, 24 rows between lateral veins. Spores with spines.
Epiphytic or epilithic on dry or muddy rocks in deciduous forests at low to medium elevations, fairly common on limestone rocks; 1001300(1700) m. Guangdong, Guangxi, Guizhou, Hainan, Yunnan [India, Indochina [??countries], Malaysia, Thailand].
Drynaria bonii has been wrongly identified as Drynaria sparsisora (Desvaux) T. Moore (Polypodium sparsisorum Desvaux; P. linnaei Bory; Drynaria linnaei (Bory) Beddome), a species of Southeast Asia and Australia, which differs by the basal fronds more deeply lobed, up to 1/3, and rhizome scales with a very long acumen.
槲蕨 hu jue
Drynaria fortunei (Kunze ex Mettenius) J. Smith (1857) [??nom illeg.], not T. Moore (1855); Polypodium fortunei Kunze ex Mettenius (Farngatt. 1, Polyp. 121, t. 3: 4245. 1857 [??1856, not (T. Moore) Lowe 1856]).
Rhizome shortly creeping, 12 cm in diam.; scales peltate, 712 Χ 0.81.5 mm, margin dentate; fronds dimorphic, glabrous; basal fronds overlapping, sessile, (2)59 Χ (2) 37 cm, base cordate, shallowly lobed to 1/3, abaxially sparsely hairy; foliage fronds stalked, stipe 47(13) cm, conspicuously winged; lamina pinnatifid up to 25 mm from costa, 2045 Χ 1015(20) cm, apex aborted or not; pinnae 713 pairs, slightly ascending, lanceolate, 610 Χ (1.5) 23 cm, margin obscurely dentate, apex obtuse or acute. Sori in 24 rows between costa and margin, one row between lateral veins, paraphyses many, glandular. Spores with globules.
Epiphytic or epilithic, often on limestone, or on buildings; 1001800 m. Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan [India (Assam), Indochina [??countries], N Thailand].
The rhizome of Drynaria roosii is used in traditional Chinese medicine as a medicinal herb, also known as a kind of Gu sui bu, [normally derived from?] Davallia.
栎叶槲蕨 le ye hu jue
Polypodium quercifolium Linnaeus, Sp. Pl. 2: 1087. 1753.
Rhizome shortly creeping, 23 cm in diam. or more; scales spreading, blackish brown, linear, 620 Χ 0.51 mm, pseudopeltate or peltate, toward apex strongly dentate, apex long, narrow, acute; fronds dimorphic; basal fronds sessile, 1550 Χ 1030 cm, shallowly lobed; foliage fronds stalked, stipe up to 30 cm, not or inconspicuously winged; lamina pinnatifid up to 25 mm from costa, 40100 Χ ca. 40 cm, apex aborted; pinnae broadly lanceolate, 1525 Χ 23.5 cm, apex acute or acuminate. Sori in 2 rows between costa and margin, regular or irregular in two rows between lateral veins, slightly sunken. Spores with acuminate spines.
Epiphytic, spirally climbing, occasionaly epilithic or terrestrial, in various types of primary and secondary forest, plantations; sea level up to 1000 m. Hainan [E Himalaya, Sri Lanka, Indochina to SE Asia[list countries]; Australia].
小槲蕨 xiao hu jue
Pleopeltis parishii Beddome, Ferns Brit. Ind. t. 125. 1866; Drynaria mutilata H. Christ.
Rhizome rather long-creeping, terete, ca. 1 cm in diam.; scales appressed, gray-brown, dark at middle, peltate, 47 Χ 11.8 mm, margin gray, long-ciliate; fronds monomorphic, glabrous; basal fronds absent; foliage fronds stalked; stipe 510 cm, conspicuously winged to very base; lamina pinnatifid up to ca. 2 mm from costa, ca. 25 Χ 20 cm, apex aborted; pinnae 59(11) pairs, spreading or slightly ascending, oblanceolate, 410 Χ 11.5(2) cm, upper 1/3 broadest, gradually narrowed toward base, margin subentire, apex acute or acuminate; venation prominent, 25 rows of areolae between lateral veins, with free included veinlets. Sori in one row between costa and margin, close to costa, only one sorus between lateral veins, obviously sunken and prominent on adaxial surface.
Epiphytic or epilithic, often on tree trunks in moss forests; 5001600 m. S and SE Yunnan [Indochina [??list countries], Myanmar, Thailand].
石莲姜槲蕨 shi liang jiang hu jue
Polypodium propinquum Wallich ex Mettenius, Abh. Senckenberg. Naturf. Ges. 2: 120. 1857[??1856]; Drynaria propinqua var. mesosora H. Christ; Phymatodes propinqua C. Presl, nom. nud.[??]
Rhizome rather long-creeping, terete, 12 cm in diam.; scales appressed, brown, peltate, 36 Χ 11.5 mm, margin dentate; fronds dimorphic, glabrous; basal fronds rounded or ovate, 1020 Χ 718 cm, pinnatifid up to 2/3 or more, margin irregularly dentate; foliage fronds stalked, stipe 820(25) cm, slightly winged; lamina pinnatifid up to ca. 2 mm from costa, (20)3050(60) Χ (12)2030 cm, apex not aborted; pinnae 715 pairs, margin dentate, apex acute. Sori in one regular row between costa and margin, close to costa. Spores verrucate and with spines.
Epiphytic or epilithic; 5001900(2800) m. Guangxi, Guizhou, Sichuan, Xizang, Yunnan [Bhutan, N India, Indochina [??countries], Myanmar, Nepal, Thailand].
毛槲蕨 mao hu jue
Drynaria costulisora Ching & S. K. Wu; D. rivalis (Mettenius ex Baker) H. Christ; D. tibetica Ching & S. K. Wu; Polypodium rivale Mettenius ex Baker.
Rhizome creeping, 0.51(2) cm in diam.; scales soft, tufted, basifixed, 511 Χ 0.51.2 mm, margin biserrate; fronds dimorphic, hairy; basal fronds sessile, 715 Χ 37 cm, base cordate, auriculate, pinnatilobed up to 2/3 or more, lobes 813 pairs, margin entire, ciliate at young stage, glabrous on both sides, lower side of costa slightly hairy and with small scales at base; foliage fronds stalked, stipe 13(13) cm, conspicuously winged; lamina pinnatifid up to 02 mm from costa, 2040(50) Χ 712(15) cm, apex not aborted; pinnae 1518 pairs, spreading, lanceolate, 38 Χ 11.5(2) cm, margin entire, densely ciliate, apex obtuse; abaxial side along costa with small scales. Sori in one straight row between costa and margin, one row between lateral veins. Spores with spines.
Epilithic on limestone rocks in Quercus forests, or on tree trunks in mixed forests; 27003400 m. Xizang, NW Yunnan [Bhutan, N India, Nepal].
秦岭槲蕨 qin ling hu jue
Polypodium baronii H. Christ, Nuovo Giorn. Bot. Ital., n.s., 4(1): 100. 1897, not Baker (1887); Drynaria reducta H. Christ; D. sinica Diels; D. sinica var. intermedia Ching & S. K. Wu.
Rhizome 12 cm in diam., with many old persistent stipes, creeping; scales soft, tufted, basifixed, 411 Χ 0.51.5 mm, base auriculate, margin biserrate; fronds dimorphic, or sometimes basal fronds absent; basal fronds sessile or with very short stipe, oblong-lanceolate, 515 Χ 36 cm, base narrowed, not cordate, not auriculate, pinnatilobed up to 2/3 or more, lobes 1012(20) pairs, margin slightly dentate; foliage fronds stalked, stipe 210 cm, conspicuously winged; lamina pinnatifid, 2250 Χ 712 cm, apex often aborted; pinnae 1625(30) pairs, spreading, lanceolate, 47 Χ 0.51.2 cm, margin dentate, glabrous or sparsely ciliate, apex acute; lamina sparsely hairy on both surfaces, especially along costa and veins. Sori in one straight row between costa and margin, close to costa, one row between lateral veins. Spores verrucate and with spines.
· Often terrestrial, or epilithic, rarely epiphytic on tree trunks; 13003800 m. Gansu, Qinghai, Shaanxi, Shanxi, Sichuan, E Xizang, NW Yunnan.
川滇槲蕨 chuan dian hu jue
Drynaria rivalis (Mettenius ex Baker) H. Christ var. yunnanensis H. Christ, Bull. Herb. Boissier 7(1): 6. 1899.
Rhizome 12 cm in diam., with many old persistent stipes, creeping; scales soft, tufted, basifixed, 410 Χ 0.51 mm, base auriculate, margin biserrate; fronds dimorphic; basal fronds sessile, ovate to oblong, 613(-17) Χ 410 cm, base auriculate, pinnatilobed up to 2/3 or more, lobes 57 pairs, margin slightly dentate; foliage fronds stalked, stipe 39 cm, winged; lamina pinnatifid, 2545 Χ 1218 cm, apex aborted or not; pinnae 713(17) pairs, spreading, lanceolate, (5)7.512(14) Χ 1.52(3.5) cm, margin shallowly dentate, glabrous or with few hairs, apex acute; lamina sparsely hairy on both surfaces, especially along costa and veins. Sori in one straight row between costa and margin, close to costa, one row between lateral veins. Sporangia often with glandular hairs. Spores verrucate.
Epiphytic or epilithic; 10001900(4200) m. S Gansu, Qinghai, Shaanxi, Sichuan, E Xizang, NW Yunnan [Bhutan, Myanmar].
Zhang Xianchun (张宪春); Hans P. Nooteboom
雨蕨属 yu jue shu
Rhizome shortly creeping. Roots restricted to ventral side; scales not hairy but with marginal setae at least in distal part, not toothed, smooth adaxially, basifixed with cordate base and much overlapping basal lobes. Fronds monomorphic; stipe articulate to phyllopodia, grooved, glabrous or with few scales; lamina compound, 3- or 4-pinnate toward base and in middle part, toward base deltoid and broadest or elongate, often narrowed, glabrous; pinnae linear-triangular; pinnules of at least larger pinnae anadromous; pinnules or pinna lobes linear-oblong; ultimate segments or lobes obtuse or acute without a pronounced tooth. Rachis adaxially grooved. Lamina axes glabrous. Veins in ultimate lobes simple, not reaching margin; false veins not present. Sori exindusiate, frequently single on a segment, facing midveins at bending point.
One species: E Asia.
Ching (Acta Phytotax. Sin. 16(3): 4; 16(4): 32. 1978) pointed out that the resemblance of Gymnogrammitis to the Davalliaceae, where this genus was previously placed, is superficial and suggested that it belongs in the Polypodiaceae. Recent molecular phylogeny has confirmed this and shown that the closest relationship is to the Selligueoid lineage of the Polypodiaceae.
雨蕨 yu jue
Polypodium dareiforme Hooker, Sec. Cent. Ferns, t. 24. 1860, [dareaeforme]; Araiostegia dareiformis (Hooker) Copeland[??okay]; Davallia dareiformis (Hooker) Levinge ex C. B. Clarke; Leucostegia dareiformis (Hooker) Beddomme; P. dareiformioides Ching.
Rhizome short, without scales 45 mm in diam.[meaning 45 mm in diam. (excluding scales)?]; scales light brown, without pale border, 210 mm, evenly narrowed toward apex. Stipe dark brown, 415 cm; lamina 3- or 4-pinnate toward base and in middle part, triangular and broadest toward base, or elongate, often narrowed toward base, 737 Χ 722 cm, glabrous; longest petiolules 26 mm; longest pinnae 4.520 Χ 1.56 cm; longest pinnules or pinna lobes 1540 Χ 625 mm; ultimate pinnules linear-oblong, lobed almost to midrib; ultimate segments 0.55 Χ 0.51 mm.
Epiphytic on mossy tree trunks, epilithic in evergreen forests; 12002600 m. Guangxi, Guizhou, Hainan, Hunan, Yunnan [Bhutan, India, Myanmar, Nepal, Thailand].