key to the genera of tribe Lepisoreae:
The characters of the tribe as defined by Hennipman et al. (1990) include a usually entire lamina, clathrate stem scales and spores with a Belvisia-type exospore and thin perispore. However, the ultrastructure of the spores was studied only for species belonging to Lepisorus (van Uffelen, 1997 [??page citation]).
[??Key 1 and Key 2]
1a...... Fronds prominently pedately to palmately divided ................................................................ 20. Neocheiropteris
1b..... Fronds simple, pinnately divided, or pinnate, rarely shallowly palmately lobed (Lepisorus waltonii).
2a...... Plants climbing; rhizome scales bearing a tuft of long, stiff, needlelike, reddish brown hairs abaxially near their center 29. Tricholepidium
2b..... Plants epiphytic, terrestrial, or climbing; rhizome scales glabrous, occasionally bearing a tuft of short, soft, brown hairs adaxially at point of attachment.
3a...... Sori scattered or arranged into ± straight lines, never merging into coenosori.
4a. Plants terrestrial; sori arranged into (1 or)24 lines on each side of midrib ................ 22. Neolepisorus
4b..... Plants climbing; sori scattered over lamina, not in defined lines ....................... 30. Lepidomicrosorium
3b..... Sori arranged into two lines, one on each side of costa or costule, discrete or merging into longitudinal coenosori.
5a. Lamina with conspicuous reddish linear scales on both surfaces ................................... 28. Caobangia
5b..... Lamina with scales absent or short and inconspicuous.
6a...... Plants epiphytic or terrestrial; rhizomes short- to long-creeping; sori discrete, sometimes coenosoral; always with clathrate, peltate paraphyses ................................................................................................ 23. Lepisorus
6b..... Plants climbing; rhizomes long and slender; sori discrete or coenosoral, with or rarely without paraphyses, these clathrate, peltate, of thick texture and irregular margin ........................................... 26. Lemmaphyllum
[??] .. Lepidogrammitis
[genera missing: Goniophlebium, Metapolypodium, Polypodiastrum, Polypodiodes, Tricholepidium, Neolepisorus ]
1a...... Fronds pedately to palmately divided ....................................................................................... 20. Neocheiropteris
1b..... Fronds simple or pinnately compound to divided, occasionally hastate to very unequally 3-lobed.
2a...... Lateral pinnae articulate to rachis ........................................................................................... 16. Goniophlebium
2b..... Lateral pinnae not articulate at base.
3a...... Veins free, without areoles along costa.
[3a.... Veinlets pinnate; scales on rhizome thick, not clathrate ................................................. 36. Polypodium]
3b..... Veinlets forked; scales on rhizome thin, pellucid, clathrate .................................... 17. Metapolypodium
3b..... Veins anastomosing to form areoles along costa or rachis.
4a. Lamina truly pinnate with lower portion of rachis terete ........................................... 18. Polypodiastrum
4b..... Lamina pinnatipartite, rachis winged throughout .......................................................... 19. Polypodiodes
棱脉蕨属 len mai jue shu
Polypodium sect. Goniophlebium Blume, Fl. Javae Filic. 132. 1830; P. subgen. Goniophlebium (Blume) C. Christensen; Schellolepis J. Smith.
Plants epiphytic, medium-sized. Rhizome long-creeping, densely scaly; scales brown to dark brown, pseudopeltate, lanceolate, clathrate, base broad, apex narrowly acuminate. Fronds remote, monomorphic; stipe straw-colored, long, glabrous; lamina imparipinnate, oblong in outline; lateral pinnae 2040 pairs, far apart, at least basal ones articulate, lanceolate or linear-lanceolate, herbaceous, glabrous or pubescent, rarely scaly, margins toothed or incised; veins anastomosing to form 2 or 3 rows of areoles on either side of costa, each areole with a simple included veinlet, which start from basiscopic lateral vein, outer veinlets free. Sori round, in one row on either side of costa, borne on simple included veinlets in costal areoles, ± immersed on abaxial surface and raised on adaxial surface. Paraphyses present only when young, peltate, clathrate, toothed at margin. Sporangia long-stalked, annulus with 12 indurate cells. Spores yellow, ellipsoidal, with obscure aperture, exospore with tuberculate surface, perispore pellucid. Χ = 37.
About 20 species: mainly in tropical Asia; two species in China (Hainan and Yunnan).
1a...... Rhizome green or brown; scales ovate-lanceolate, margins ciliate; lateral pinnae with 45 mm stalks, base cuneate 1. G. persicifolium
1b..... Rhizome glaucous underneath scales; scales linear-lanceolate, margins entire; lateral pinnae sessile, base cordate or subcordate, auriculate on both sides or on acroscopic side only ...................................... 2. G. subauriculatum
棱脉蕨 len mai jue
Polypodium persicifolium Desvaux, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 5: 316. 1811; Goniophlebium grandidens (Mettenius) Fιe; G. ponapense Copeland; G. rotense Hosokawa; P. colpothrix Kunze; P. grandidens Kunze ex Mettenius; P. integrius Copeland[??integriore]; P. koningsbergeri Alderwerelt; P. persicifolium var. mettenii Rosenstock; P. phlebodioides Copeland; Schellolepis persicifolia (Desvaux) Pichi Sermolli.
Rhizome green or brown, 57 mm in diam., densely scaly; scales dark brown, ovate-lanceolate, 34 Χ 1.52 mm, margin toothed. Stipe straw-colored or brownish, 3040 cm, 34 mm in diam., densely scaly at base, distally minutely scaly or glabrescent upward. Lamina broadly lanceolate in outline, up to 150 Χ 40 cm; lateral pinnae 1030 pairs, ascending, 45 cm apart, ± falcate, linear-lanceolate, up to 20 Χ 2.5 cm, cuneate at base, with 57 mm stalks, margin toothed and cartilaginous, apex caudate-acuminate; pinnae gradually smaller distally, terminal pinna same as lateral pinnae, up to 15 cm; lamina herbaceous or subleathery, green, glabrous. Sori round, borne on simple included veinlets in costal areoles, in one row on each side of costa, medial or costular, distinctly immersed on abaxial surface and raised on adaxial surface.
Epiphytic on tree trunks; 7001000 m. Hainan (Baotin, Diaoluo Shan, Jianfeng Ling) [SE India, Indonesia, Malaysia, New Guinea, Philippines, Thailand, Vietnam; Pacific islands].
Beddome (Ferns Brit. Ind. 79. pl. 79. 1865) treated material of this taxon as Goniophlebium cuspidatum (D. Don) C. Presl.
穴果棱脉蕨 xue guo len mai jue
Polypodium subauriculatum Blume, Enum. Pl. Javae 2: 133. 1828; Goniophlebium molle Beddome; Marginaria subauriculata (Blume) Nakai ex H. Itτ; Polypodiastrum molle (Beddome) Ching; Polypodium beddomei Baker; Schellolepis subauriculata (Blume) J. Smith.
Rhizome 68 mm diam., glaucous underneath scales, densely scaly at least at apex; scales dark brown, narrowly lanceolate, ca. 5 Χ 0.7 mm, margin toothed. Stipe straw-colored or brown, 2535 cm, base densely scaly, distally minutely scaly or glabrescent; rachis pale brown, minutely scaly throughout; lamina lanceolate in outline, 80100 Χ 3040 cm; lateral pinnae 3040 pairs, sessile, linear, basal pinnae usually slightly shorter than next above, deflexed or spreading; middle pinnae subopposite, spreading or slightly ascending, straight or slightly falcate, 1420 Χ ca. 2 cm, base subcordate or truncate, rounded auriculate on both sides, gradually narrowing from base to long-attenuate apex, margin serrate, upper pinnae progressively smaller; terminal pinna 510 cm, basal portion irregularly lobed; lamina herbaceous or thinly leathery, green, glabrous or pubescent when young. Sori rounded, at tip of simple included veinlets in costal areoles, in one row on each side of costa, distinctly immersed on abaxial surface and raised on adaxial surface.
Epiphytic on tree trunks; 5001300 m. Yunnan (Jinhong, Mengla). [Indonesia, Malaya, New Guinea, Philippines, Thailand, Vietnam; NE Australia, Pacific islands].
篦齿蕨属 bi chi jue shu
Ferns, epiphytic, medium-sized. Rhizome long-creeping, densely covered with scales; scales dark brown, lanceolate, base broad and reddish brown ciliate, acuminate at apex. Fronds remote. Stipe stramineous, articulate to rhizome. Lamina narrowly lanceolate, deeply pinnatifid or pinnatisect at lower part. Lateral lobes more than 30 pairs, narrowly lanceolate, spreading or slightly deflexed at lower pairs. Veins free, veinlets forked. Sori round, in a single row on each side of costa, paraphyses absent.
Monotypic genus in Asia, from SW China to Indo-China.
篦齿蕨 bi chi jue
Polypodium manmeiense H. Christ, Bull. Herb. Boissier 6: 870. 1898; Goniophlebium manmeiense (H. Christ) Rφdl-Linder; Metapolypodium kingpingense Ching & W. M. Chu; Polypodiodes manmeiense (H. Christ) Fraser-Jenkins; Polypodium pseudodimidiatum H. Christ; P. scalare H. Christ; P. simulans Baker.
Rhizome long-creeping, 23 mm in diam., densely covered with scales; scales dark brown, lanceolate, broad at base, remotely ciliate at margin, acuminate at apex. Fronds remote. Stipe straw-colored or brown at base, 812 cm, densely scaly at base, glabrous upward. Lamina narrowly lanceolate, 2030 Χ 57 cm, deeply pinnatifid, pinnae usually decurrent to next lobes by very narrow wings of rachis in lower part, with rather abruptly narrowing acuminate apex. Lateral lobes 2030 pairs, narrowly lanceolate, spreading except base with one pair deflexed and slightly shortened, incised-undulate at margins, acute to obtuse at apex. Rachis and costa straw-colored. Veins free, veinlets forked, terminally veinlets ending in elliptic hydathodes inside margin of lobes. Texture herbaceous, green, glabrous. Sori round, in one row on each side of costa, medial, superficial or immersed in cavities on abaxial surface.
Epiphytic on tree trunks or on rocks; 10002500 m. Guizhou, Sichuan, Yunnan [Cambodia, NE India, Laos, N Myanmar, Thailand, Vietnam].
拟水龙骨属 ni shui long gu shu
Ferns, medium-sized, epiphytic. Rhizome long-creeping, densely scaly; scales brown to dark brown, lanceolate, clathrate. Fronds remote. Stipe articulate to rhizome at base. Lamina pinnate. Pinnae ca. 10 pairs, sessile, lanceolate to narrowly lanceolate, upper pinnae usually decurrent at base. Veins anastomosing to form l3 rows of areoles on each side of costa, each costal areole containing an included free veinlet. Texture herbaceous, glabrous or minutely scaly on abaxial side. Sori round, terminal on included veinlets of costal areoles, in one row on each side of costa; sporangia long stalked, annulus with about 1220 indurate cells; receptacular scalelike, clathrate, fugacious; spores ellipsoidal, surface coarsely verrucate. x=37.
About eight species: tropical and subtropical Asia and Oceania; three species in China.
1a...... Rhizome densely covered with scales; lateral pinnae deflexed or spreading, decurrent at base; costa straw-colored, veinlets invisible ................................................................................................................................................ 1. P. dielseanum
1b..... Rhizome sparsely covered with scales and with whitish bloom; lateral pinnae ascending, rounded or cordate at base; costa brown, veinlets visible.
2a...... Lateral pinnae rounded at base, not overlapping rachis ............................................................... 2. P. argutum
2b..... Lateral pinnae cordate at base, overlapping rachis ................................................................ 3. P. mengtzeense
川拟水龙骨 chuan ni shui long gu
Polypodium dielseanum C. Christensen, Index Filic. 9: 522. 1906; P. leuconeuron Diels (Bot. Jahrb. Syst. 29: 203. 1900), not (Fιe) H. Christ (1900); Goniophlebium dielseanum (C. Christensen) Rφdl-Linder; P. meyii H. Christ, [meyi]; P. wilsonii H. Christ.
Rhizome long-creeping, ca. 5 mm in diam., densely scaly; scales brown, ovate-lanceolate, minutely denticulate at margins, acuminate at base. Fronds remote. Stipe straw-colored, 2030 cm, densely scaly at base, upward glabrous, rachis and lower part of costa pubescent on both surfaces; lamina pinnate, ovate-lanceolate in outline, 4060 Χ 1525 cm; lateral pinnae 2030 pairs, basal pinna deflexed, upper pinnae spreading, linear-lanceolate, 1015 Χ 0.8l.2 cm, base adnate and somewhat dilated, margin crenate-serrate, apex acuminate or obtuse; veins anastomosing to form one row of areoles on each side of costa with included veinlet. Texture herbaceous, green, abaxial surface with sparse brown ovate-lanceolate scales. Sori round, in one row on each side of costa, terminal on included veinlets, medial.
Epiphytic on tree trunks or on rocks; 15002010 m. Sichuan, Guizhou, Yunnan [NE India].
尖齿拟水龙骨 jian chi ni shui long gu
Rhizome long-creeping, 34 mm diam., covered with sparse scales and whitish bloom; scales brown, ovate-lanceolate, peltate at broad base, sparsely toothed at margin, acuminate at apex. Fronds remote. Stipes straw-colored, 1015 cm, scaly at base, glabrous upward. Lamina ovate lanceolate in outline, 4050 Χ 1525 cm. Lateral pinnae 1520 pairs, sessile, ovate-lanceolate, 1015 Χ 1.52.5 cm, base rounded, margin serrate or denticulate, apex acuminate; terminal pinna distinct; veins anastomosing to form 2 or 3 areoles on each side of costa, each costal areole containing a simple free included veinlet, other free veinlet ending inside margin of lobes, visible on both surfaces. Lamina herbaceous, abaxially yellowish green, adaxially dark green, both surfaces glabrous except for minute scales abaxially. Sori round, terminal on included veinlets of costal areoles, in one row on each side of costa, medial or slightly closer to costa, superficial.
Epiphytic on tree trunks or on rocks; 23002700 m. Guangxi, Guizhou, Xizang, Yunnan [Bhutan, NE India, Myanmar, Thailand, Nepal].
The epithet argutum was first used by Wallich (Cat. No. 308. l828, nom. nud.).
1a. Pinnae 1520 pairs, ovate-lanceolate, margin crenate-serrate; sori medial ................................. 2a. var. argutum
1b. Pinnae fewer and narrower, margin denticulate; sori closer to costa ........................................ 2b. var. angustum
尖齿拟水龙骨(原变种) jian chi ni shui long gu (yuan bian zhong)
Polypodium argutum Wallich ex Hooker, Sp. Fil. 5: 5: 32. 1864 [??1863]; Goniophlebium argutum (Wallich ex Hooker) Beddome [??J. Smith ex Hooker (1842)]; Schellolepis arguta (Wallich ex Hooker) J. Smith.
Pinnae 1520 pairs, ovate-lanceolate, margin crenate-serrate. Sori medial.
Epiphytic on tree trunks or on rocks; 23002700 m. Guangxi, Guizhou, Xizang, Yunnan [Bhutan, NE India, Myanmar, Thailand, Nepal].
狭羽拟水龙骨 xia yu ni shui long gu
Pinnae fewer and narrower, margin denticulate. Sori closer to costa.
* Epiphytic on tree trunks; 23002400 m. Xizang.
蒙自拟水龙骨 meng zi ni shui long gu
Rhizome long-creeping, 45 mm diam., covered with whitish bloom and sparse scales; scales dark brown, ovate-lanceolate, peltate at broad base, irregularly toothed at margin, acuminate at apex. Fronds remote. Stipe pale straw-colored, 1020 cm, scaly at base, glabrous upward. Lamina ovate-lanceolate, pinnate with distinct terminal pinna, 5070 Χ 1520 cm. Lateral pinnae l525 pairs, subopposite, sessile, lanceolate, 1015 Χ 1.52 cm, obliquely cordate at base, ± rounded auriculate, overlapping rachis, serrate at margin, acuminate at apex. Veins anastomosing to form 1 or 2 rows of areoles on each side of costa, each costal areole containing a simple free veinlet, other veinlets free, ending inside margin of teeth. Texture herbaceous, yellowish green, glabrous on both surfaces except sparsely scaly on abaxial side of rachis and costa. Sori round, in one row on each side of costa, terminal on included veinlets, medial or slightly closer to costa.
Epiphytic on tree trunks or on rocks, l5002500 m. Guangdong, Guangxi, Guizhou, Taiwan, Yunnan [NE India, Laos, Nepal, Philippines, Thailand, Vietnam].
Material of this species (Polypodiastrum mengtzeense) was treated by Wu (Sunyatsenia 3: 266, pl. 124. 1932) as Polypodium argutum Wallich ex Hooker.
水龙骨属 shui long gu shu
Ferns, medium-sized. Rhizome long-creeping, mostly densely scaly, with whitish bloom when scales sparse; scales brown to dark brown, lanceolate to ovate-lanceolate, clathrate, peltate at broad base, margin entire, denticulate, or ciliate, apex acuminate or hair-shaped. Fronds remote, articulate to rhizome; stipe straw-colored or brown, scaly at base; lamina deeply pinnatisect, oblong-lanceolate in outline; segments 1060 pairs, narrowly lanceolate, bases confluent through narrowly winged rachis, rarely lowermost free, margin toothed or serrulate, apex acuminate. Veins usually anastomosing to form a row of very narrow areoles on each side of costa and 1 or 2 rows of much broader areoles along each side of costules, latter each including a free veinlet. Lamina herbaceous, glabrous or pubescent, usually minutely scaly abaxially. Sori round, in single rows on each side of costules, each at end of included veinlet, superficial or immersed, paraphyses soon lost; sporangia long-stalked, annulus with about 1219 indurate cells; spores ellipsoidal, surface coasely verrucate. Χ = 37.
About 17 species: mostly confined to tropical and subtropical Asia; 12 species in China (three endemic).
Christenhusz et al. (2010 [??page citation]) treat Polypodiodes as a synonym of Goniophlebium.
1a...... Rhizome covered with whitish bloom and sparse scales or scales only present on very young part; segment margins entire.
2a...... Lamina abaxially glabrous.
3a...... Rhizome
scales usually absent; lamina adaxially pubescent only along rachis and costae
........................................................................................................................................................ l.
P. formosana
3b..... Rhizome scales black, linear-subulate and with a broad brown base; lamina adaxially more uniformly pubescent 2. P. raishanensis
2b..... Lamina abaxially pubescent.
4a...... Rachis wings 23 mm wide; segments 0.51 cm wide .............................................................. 3. P. niponica
4b..... Rachis wings ca. 5 mm wide; segments 1.21.4 cm wide ............................................................... 4. P. wattii
1b..... Rhizome usually densely scaly, without whitish bloom; segment margins usually incised to serrate.
5a...... Rhizome scales light brown, 89 mm ............................................................................................... 5. P. bourretii
5b..... Rhizome scales brown to dark brown or black, 25 mm.
6a...... Rhizome scales linear-subulate, relatively sparse; segment margins entire ........ 5. P. transpianensis[??]
6b. Rhizome scales wider, usually concealing rhizome; segment margins incised to serrate.
7a. Veinlets along rachis free, not forming areoles; upper stipe and rachis chestnut .. 6. P. microrhizoma
7b..... Veinlets along rachis anastomosing to form row of narrow areoles on each side; stipe and rachis mostly straw-colored.
8a...... Lamina linear-lanceolate in outline; lateral segments up to 4060 pairs.
9a...... Scales black and subulate at upper part, brown at broad base, margin ciliate; fronds subglabrous 7. P. lachnopus
9b..... Scales brown, lanceolate, margin irregularly denticulate; fronds pubescent on both sides .. 8. P. pseudolachnopus
8b..... Lamina ovate-lanceolate in outline; lateral segments usually 1020 pairs.
10a.... Segment margins densely double-serrate.
11a.... Rhizome scales ovate-lanceolate, brown, margin entire .................................... 9. P. subamoena
11b... Rhizome
scales narrowly lanceolate, black, apex hairlike, margin denticulate
.................................................................................................................................. 10.
P. hendersonii
10b... Segment margins incised or incised-serrate.
12a.... Rhizome 34 mm in diam., scales black; lamina segments 48 mm wide; sori close to costa 11. P. chinensis
12b... Rhizome
57 mm in diam., scales brown, lamina segments 1518 wide; sori medial
......................................................................................................................................... 12.
P. amoena
台湾水龙骨 tai wan shui long gu
Polypodium formosanum Baker, J. Bot. 23: 105. 1885 [not P. formosum Raddi (1825)]; Goniophlebium formosanum (Baker) Rφdl-Linder; Marginaria formosana (Baker) Nakai ex H. Itτ; P. liukiuense H. Christ.
Rhizome long-creeping, ca. 4 mm in diam., covered with whitish bloom; scales fugacious, only present on very young parts, brown, ovate-lanceolate, entire at margin, acuminate at apex. Fronds remote. Stipe straw-colored, 1520 cm, glabrous. Lamina oblong, 3050 Χ 1015 cm, cordate at base, pinnatipartite, acuminate or caudate at apex. Segments 2030 pairs, narrowly lanceolate, 58 Χ 0.81 cm, entire at margin, acuminate at apex; lowest segments usually deflexed. Veins anastomosing to form 1 or 2 rows of areoles on each side of costa with included veinlets. Texture herbaceous, green, abaxial surface glabrous or subglabrous, adaxial surface pubescent, hairs on rachis and costa more dense. Sori round, terminal on included veinlets, in one row on each side of costa, medial.
Epiphytic on tree trunks or on rocks; 2001200 m. Fujian, Taiwan [Japan].
大叶水龙骨 da ye shui long gu
Polypodium raishanense Rosenstock, Hedwigia 56: 346. 1915; Marginaria pseudoformosana Tagawa; M. raishanensis (Rosenstock) Nakai ex H. Itτ.
Rhizome long-creeping, ca. 3 mm in diam., covered with whitish bloom; scales sparse, black, linear-subulate and with a broad brown base. Fronds remote. Stipe and rachis glossy, turning brown when mature. Lamina oblong, 3055 Χ 1015 cm, pinnatipartite. Segments spreading, 4.58 cm, adnate to rachis by a narrow wing, margin entire, abaxial surface glabrous, pubescent on margin and on adaxial surface; lower segments shortened and deflexed. Veins anastomosing to form 1 or 2 rows of areoles on each side of costa with included veinlets. Texture herbaceous. Sori round, small, closely spaced, borne near costa in costal areolae at end of free veinlets.
* Epiphytic on tree trunks or on rocks; 2001200 m. Taiwan.
日本水龙骨 ri ben shui long gu
Polypodium niponicum Mettenius, Ann. Mus. Bot. Lugduno-Batavi 2: 222. 1866; Marginaria niponica (Mettenius) Nakai ex H. Itτ; M. transpianensis (Yamamoto) H. Itτ; Polypodiodes transpianensis (Yamamoto) Saiki; Polypodium bodinieri H. Christ; P. longkyense Rosenstock; P. silvestrii H. Christ; P. transpianense Yamamoto.
Rhizome long-creeping, gray-green, ca. 5 mm in diam., sparsely covered with whitish bloom and scales; scales dark brown, narrowly lanceolate, ovate at base, denticulate at margin, acuminate at apex. Stipe straw-colored, 515 cm, pubescent. Lamina oblong-lanceolate, 3040 Χ 1013 cm, pinnatipartite, cordate at base, pinnatilobate-acuminate at apex. Segments 1525 pairs, linear-oblong, 35 Χ 0.51 cm, entire at margin, acute or obtuse at apex; lowest ones often subfree, slightly shortened and deflexed. Veins anastomosing to form one row of areoles at each side of rachis and each side of costa, veinlets invisible. Texture herbaceous, gray-green, pubescent on both surfaces or more densely so abaxially with short, whitish, 2- or 3-celled, soft hairs. Sori round, in one row on each side of costa, terminal on included veinlets in costal areoles, nearer to costa than to margin.
Epiphytic on tree trunks or on rocks; 3001600 m. Anhui, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [NE India, Japan, Vietnam].
Material of Polypodiodes niponica from Taiwan with the rhizomes lacking the whitish bloom and with very narrow scales has been separated as Polypodiodes transpianensis.
光茎水龙骨 guang jing shui long gu
Polypodium niponicum Mettenius var. wattii Beddome, J. Bot. 26: 235. 1888; Goniophlebium niponicum var. wattii (Beddome) Beddome; Polypodiodes niponica (Mettenius) Ching var. wattii (Beddome) W. M. Chu & S. G. Lu; Polypodium niponicum var. laevipes Franchet ex H. Christ.
Rhizome fleshy, long-creeping, 57 mm in diam., covered with whitish bloom, scales only present on growing tip or at base of stipes; scales dark brown, lanceolate, minutely serrate at margin, acuminate at apex. Stipe dark straw-colored, 515 cm, pubescent. Lamina oblong, 2030 Χ 810 cm, pinnatipartite, wings of rachis ca. 5 mm on each side, truncate to cordate at base, caudate at apex. Segments 1520 pairs, 45 Χ 1.21.4 cm, entire at margin, obtuse at apex; lowest ones subspreading to slightly deflexed. Veins anastomosing to form 2 or 3 rows of areoles on each side of costa and one row of areoles on each side of rachis, costal areoles with included free veinlets. Texture herbaceous, yellowish green or gray-green, pubescent and whitish short hairy on both surfaces or more densely so abaxially. Sori round, in one row on each side of costa, terminal on included veinlets, slightly close to midrib.
Epiphytic on tree trunks or on rocks; 13003000 m. Sichuan, Xizang, Yunnan [NE India, N Myanmar, Vietnam].
[??]
滇越水龙骨 dian yue shui long gu
Polypodium bourretii C. Christensen & Tardieu, Notul. Syst. (Paris) 8: 183. 1939; P. wangii Ching (1949), not C. Christensen (1934).
Rhizome long-creeping, ca. 4 mm diam., densely covered with scales; scales light brown, linear-subulate, clathrate, ca. 10 mm, broad at base with minutely serrate margin, upper part acuminate with entire margin, hairlike at apex. Fronds remote. Stipe straw-colored, 1015 cm, slender, glabrous. Lamina oblong-lanceolate, 2040 Χ 814 cm, cordate and slightly narrowed at base, pinnatipartite, pinnatifid-acuminate at apex. Segments 2030 pairs, linear-lanceolate, 47 cm Χ 68 mm, base broad and connected by narrow wings along midrib, remotely incised at margin, acute at apex; basal pair subdeflexed and slightly shorter than next upper pairs. Veins anastomosing to form one row of areoles on each side of costa and of rachis. Texture herbaceous, grayish green, adaxial side sparsely covered with long, appressed, septate, whitish hairs, abaxial side densely pubescent with short hairs. Sori round, in one row on each side of costa, close to costa.
Epiphytic on tree trunks or on rocks; 6001500 m. S Guizhou, SE Yunnan [Vietnam].
栗柄水龙骨 li bing shui long gu
Polypodium microrhizoma C. B. Clarke ex Baker, Syn. Fil. (Hooker & Baker), ed. 2: 511. 1874; Goniophlebium fieldingianum (Mettenius) T. Moore; G. microrhizoma (C. B. Clarke ex Baker) Beddome; Polypodium fieldingianum Kunze ex Mettenius; P. microrhizoma var. xerophyticum Mehra, nom. illeg.; P. taliense H. Christ.
Rhizome long-creeping, 23 mm in diam., densely covered with scales; scales dark brown, ovate-lanceolate, broad at base, peltate, denticulate at margin, acuminate at apex. Fronds remote. Stipe straw-colored at base, shiny and castaneous on lower side (as is rachis), 812 cm, slender, glabrous. Lamina lanceolate, 2030 Χ 58 cm, deeply pinnatifid or pinnatisect at lower portion, base slightly shortened, apex pinnatilobate-acuminate. Segments/pinnae 1530 pairs, spreading, lanceolate, 45 cm Χ 68 mm, base connected by a narrowly wing along rachis, distantly incised-serrate at margin, acute or obtuse at apex; basal ones not or very slightly deflexed, slightly shorter than next above. Veins visible, anastomosing to form one row of areoles on each side of costa, each containing a simple included veinlet, without areoles on both sides of rachis. Texture thinly papery or membranous, green, glabrous on both surfaces, rachis castaneous abaxially, straw-colored and grooved adaxially, glabrous, nearly wingless in lowest portion. Sori round or oblong, terminal on included veinlets, in one row on each side of costa, medial, ± immersed.
Terrestrial in forest understories, epiphytic on tree trunks; 23003300 m. Sichuan, Taiwan, Xizang, Yunnan [Bhutan, N India, N Myanmar, Nepal, N Thailand].
DeVol and Kuo (Fl. Taiwan 1: 202. 1975) treated material of Polypodiodes microrhizoma as Polypodium atkinsonii C. Christensen.
濑水龙骨 lai shui long gu
Polypodium lachnopus Wallich ex Hooker, Hookers Icon. Pl.: t. 952. 1854; Goniophlebium lachnopus (Hooker) J. Smith; Polypodiodes lachnopus var. xerophyticum Mehra, nom. illeg.; Schellolepis lachnopus (Hooker) J. Smith.
Rhizome long-creeping, 56 mm in diam, densely covered with scales; scales broad and brown at base, ciliate at margin, upper part black, subulate, entire at margin. Fronds remote. Stipe straw-colored, 58 cm, glabrous. Lamina linear-lanceolate, 4060 Χ 57 cm, cordate at base, pinnatipartite, acuminate at apex. Segments 4050 pairs, lanceolate, 23 cm Χ 57 mm, toothed at margin, acute or obtuse at apex; basal one or two pairs slightly shortened and deflexed. Veins reticulate to form one row of narrow areoles along each side of rachis and one row of large areoles along each side of costa, each areole containing a simple included veinlet, other veins free. Texture papery or herbaceous, green, glabrous or subglabrous except rachis covered with hairs, sparsely scaly on abaxial surface. Sori round, terminal on included veinlets, in one row on each side of costa, medial.
Epiphytic on tree trunks; 17002000 m. Sichuan, Xizang, Yunnan [Bhutan, N India, Nepal].
A collection of this species (Polypodiodes lachnopus) was listed by H. Christ (Bull. Herb. Boiss. 6: 869. 1898) as Polypodium yunnanense Franchet.
假毛柄水龙骨 jia mao bing shui long gu
Rhizome long-creeping, 56 mm in diam., densely covered with scales; scales brown, lanceolate, peltate at base, irregularly denticulate at margin, acuminate at apex. Fronds remote. Stipe straw-colored, 1015 cm, sparsely pubescent. Lamina linear-lanceolate, 4050(70) Χ 68 cm, cordate at base, pinnatipartite, acuminate at apex. Segments 4060 pairs, lanceolate, 45 cm Χ 510 mm, toothed at margin, acute or obtuse at apex. Veins anastomosing to form one row of narrow areoles along each side of rachis and one row of large areoles along each side of costa, each costal areole containing a simple included veinlet. Texture papery, green, pubescent on both surfaces and scaly abaxially; rachis straw-colored, more densely hairy. Sori round, in one row on each side of costa, slightly close to costa.
* Epiphytic on tree trunks or on rocks; 18003000 m. Sichuan, Xizang, Yunnan.
Material of Polypodiodes pseudolachnopus was first identified as Polypodium lachnopus Wallich ex Hooker (Ching, Icon. Fil. Sin. 2: ad pl. 95. 1934; Shing in W. T. Wang, Vasc. Pl. Hengduan Mt. 1: 161. 1993).
假友水龙骨 jia you shui long gu
Polypodium subamoenum C. B. Clarke, Trans. Linn. Soc. London, Bot., ser. 2. 1: 550. 1880; Goniophlebium subamoenum (C. B. Clarke) Beddome.
Rhizome long-creeping, 23 mm in diam., densely covered with scales; scales brown or dark brown, ovate-lanceolate, broad at base, peltate, entire at margin, acuminate at apex. Fronds remote. Stipe stramineous, 510 cm, glabrous. Lamina ovate-lanceolate, 1520 Χ 58 cm, cordate at base, pinnatipartite, caudate at apex. Segments 1015(20) pairs, linear, 34 cm Χ 810 mm, duplicato-serrate at margin, obtuse or acute at apex; lowest pairs shortened and deflexed. Veins visible, anastomosing to form one row of areoles along rachis and 1 or 2 rows of areoles on each side of costa, costal areoles containing a simple included veinlet. Texture herbaceous, green, scaly on abaxial surface, glabrous on adaxial surface. Sori round, in one row on each side of costa, terminal on included veinlets, close to costa.
Epiphytic on tree trunks or on rocks; 24003300 m. Xizang, Yunnan [India (Sikkim), Nepal].
喜马拉雅水龙骨 xi ma la ya shui long gu
Goniophlebium hendersonii Beddome, Ferns Brit. Ind. Suppl.: 21, t. 383. 1876; Polypodiodes atkinsonii (C. Christensen) Ching; Polypodium atkinsonii C. Christensen; P. hendersonii Atkinson ex Baker (Syn. Fil. (Hooker & Baker), ed. 2: 511. 1874), not E. J. Lowe (1858).
Rhizome long-creeping, 34 mm in diam., densely covered with scales; scales black, linear-lanceolate, broad at base, peltate, toothed at margin, hairlike at apex. Fronds remote. Stipe straw-colored, 812 cm, glabrous. Lamina lanceolate, 2025 Χ 58 cm, pinnatipartite or pinnatisect at lower portion, cordate at base, acuminate at apex. Segments 2025 pairs, lanceolate, 34 Χ ca. 1 cm, serrate at margin, acuminate at apex, spreading or oblique at upper portion, lowest pair deflexed. Veins visible, anastomosing to form one row of areoles along rachis and costa, costal areoles containing a simple included veinlet. Texture papery, green, scaly on abaxial surface, glabrous on adaxial surface. Sori round, in one row on each side of costa, terminal on included veinlets, medial.
Epiphytic on tree trunks or on rocks; 20003300 m. Xizang [India (Sikkim), Nepal].
Polypodium atkinsonii was published in 1906 as a replacement name for P. hendersonii Atkinson ex Hooker & Baker (1874), not E. J. Lowe (1858). Beddome validated the epithet hendersonii under Goniophlebium and this has prioriy outside Polypodium.
中华水龙骨 zhong hua shui long gu
Polypodium subamoenum C. B. Clarke var. chinense H. Christ, Nuovo Giorn. Bot. Ital., n.s., 4: 99. 1897; Goniophlebium amoenum J. Smith var. chinense (H. Christ) Rφdl-Linder; P. amoenum Wallich ex Mettenius var. chinense (H. Christ) Ching; P. amoenum var. tonglense C. B. Clarke [??tongolense]; P. amoenum var. xerophyticum Mehra & Bir.
Rhizome long-creeping, 23 mm in diam., densely covered with scales; scales black, ovate-lanceolate, sparsely toothed at margin, acuminate at apex. Fronds remote. Stipe straw-colored, 1020 cm, glabrous. Lamina ovate-lanceolate or broadly lanceolate, 1525 Χ 710 cm, cordate at base, pinnatipartite or pinnatisect at lower portion, caudate at apex. Segments 1525 pairs, linear-lanceolate, 35 cm Χ 57 mm, serrate at margin, acuminate at apex; basal pairs slightly shortened and deflexed. Veins reticulate, costa visible, straw-colored, veinlets invisible. Texture herbaceous, subglabrous on both surfaces, scaly abaxially. Sori small, round, terminal on included veinlets, close to costa.
* Epiphytic on rocks or on tree trunks, 9002800 m. Anhui, Gansu, Guangdong, Guizhou, Hebei, Henan, Hubei, Jiangxi, Shaanxi, Shanxi, Sichuan, Taiwan, Yunnan, Zhejiang.
Rφdl-Linder (1997) treated material of this species (Polypodiodes chinensis) as a hairy variety of the following species (Polypodiodes amoena) while Fraser-Jenkins (2008 [??page citation]) suggested that it merely represents more hairy populations of P. amoena not worthy of formal recognition; he also indicates that he regards Polypodium yunnanense Franchet as part of this taxon if recognized as a distinct species and thus providing the oldest species epithet. However, the concept used here is based on a different set of characters from those discussed by Fraser Jenkins. The names Polypodium pseudo-amoenum Ching and Polypodioides pseudo-amoenum (Ching) Ching have been applied to this taxon but have never been validated. The former was first mentioned as a synonym in the protologue of Polypodium amoenum var. chinense (H. Christ) Ching.
友水龙骨 you shui long gu
Rhizome 57 mm in diam., densely covered with scales; scales dark-brown, lanceolate, base broad, margin denticulate, apex acuminate. Stipe stramineous, (5)3040 cm, glabrous. Lamina ovate-lanceolate, (11)4050 Χ (4)2025 cm, base slightly narrowed, apex acuminate. Segments 2025 pairs, lanceolate, 1013 Χ 1.52 cm, connected along rachis by a 1.54 mm wide wing, margin shallowly crenate to subserrate, apex acuminate; basal 1 or 2 pairs of segments often deflexed. Veins forming 1 or 2 rows of areoles along each side of rachis and costa. Lamina thickly papery, green, both surfaces glabrous or pubescent, abaxial surface scaly. Sori round, medial.
Epiphytic on rocks or on tree trunks; 10002500 m. Anhui, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, India, Laos, Myanmar, Nepal, Thailand, Vietnam].
The collection Purdom 32 (Tai-pei-shan) was identified by R. C. Ching as a dwarf northern form of Polypodiodes amoena. It has fronds with the stipe ca. 5 cm, lamina ca. 11 Χ 4 cm and ca. 16 pairs of segments with obtuse apices.
1a. Stipe castaneous; lamina narrowly lanceolate in outline, basal segments spreading .......... 12b. var. duclouxii
1b. Stipe straw-colored; lamina ovate-lanceolate in outline; basal 1 or 2 pairs of segments deflexed.
2a. Lamina with both surfaces glabrous ...................................................................................... 12a. var. amoena
2b. Lamina with both surfaces hairy or at least pubescent on rachis and costa ....................... 12c. var. pilosa
友水龙骨(原变种) you shui long gu (yuan bian zhong)
Polypodium amoenum Wallich ex Mettenius, Farngatt. I (Polypodium) 2(1): 131. 1857; Goniophlebium amoenum (Wallich ex Mettenius) Beddome [??J. Smith (1840)]; G. yunnanense (Franchet) Beddome; P. amoenum var. latedeltoideum H. Christ; Polypodium arisanense Hayata; P. bonatianum Brause; P. valdealatum H. Christ; P. yunnanensis Franchet [??yunnanense]; Schellolepis amoena (Wallich ex Mettenius) J. Smith.
Stipe and rachis stramineous Lamina ovate-lanceolate in outline, basal 1 or 2 pairs of segments often deflexed, both surfaces glabrous.
Epiphytic on tree trunks or on rocks; 10002500 m. Anhui, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [Bhutan, India, Laos, Myanmar, Nepal, Thailand, Vietnam].
红秆水龙骨 hong gan shui long gu
Polypodium duclouxi H. Christ, Notul. Syst. (Paris) 1: 34. 1909; Polypodium amoenum Wallich ex Mettenius var. duclouxi (H. Christ) Ching.
Stipe and abaxial surface of costae castaneous. Lamina narrowly lanceolate in outline, basal segments spreading; veinlets slender and indistinct, both surfaces glabrous.
* Epiphytic on tree trunks or on rocks; 20002500 m. Sichuan, Yunnan.
check rank, FCCL has fo. pilosa
柔毛水龙骨 rou mao shui long gu
Polypodium amoenum f. pilosum C. B. Clarke, J. Linn. Soc., Bot. 24: 417. 1888; Polypodiodes amoena f. pilosa (Rosenstock) Ching [not C. B. Clarke 1888]; Polypodium amoenum var. pilosum Rosenstock.
Stipe and rachis stramineous. Lamina ovate-lanceolate in outline, basal 1 or 2 pairs of segments often deflexed, both surfaces hairy at least on rachis and costa.
Epiphytic on tree trunks or on rocks; 15002500 m. Guizhou, Hubei, Sichuan, Xizang, Yunnan, Zhejiang [NE India, Nepal].
扇蕨属 shan jue shu
Cheiropteris H. Christ (1898), not Chiropteris J. G. Kurr ex H. G. Bronn (1858).
Plants terrestrial, medium-sized. Rhizome long-creeping, rather thick, densely scaly; scales brown, iridescent, clathrate, ovate-lanceolate, margin dentate. Stipes distant, stramineous, longer than laminae; laminae trifid, or palmately lobed, base attenuate, middle segment longer; segments lanceolate, papery, glabrous, with small brown scales when young, entire; main veins raised, small veinlets obscure. Spores verrucate.
* Two species: endemic to SW China.
DU Xin Yu & CHENG Xiao, Phylogeny of the Chinese Endemic Fern Genus Neocheiropteris (Polypodiaceae). Plant Diversity and Resources 33 (3): 261268. 2011.
1a...... Fronds palmately lobed into 810 segments .......................................................................... 1. N. palmatopedata
1b..... Fronds trifid ............................................................................................................................................. 2. N. triglossa
扇蕨 shan jue
Polypodium palmatopedatum Baker, Bull. Misc. Inform. Kew 1898: 232. 1898; Microsorum palmatopedatum (Baker) Nooteboom.
Rhizome creeping; scales slightly spreading, ovate or triangular, broadest below middle, 37 Χ 23 mm, clathrate or subclathrate, cells longitudinally rectangular, central region bearing multiseptate hairs at least when young, margin denticulate, apex acute. Fronds not or only slightly dimorphic; stipe 1040 cm; lamina pedately dissected, reniform to nearly circular in outline, 1230 Χ 2040 cm, herbaceous, abaxial surface without acicular hairs, with short glandular hairs, base cuneate, margin entire or undulate, first connecting vein forming one row of small primary costal areoles parallel to costa, other (sometimes irregularly shaped), larger areoles in one row between two veins, dichotomously branched near margin, or below middle. Sori separate, one sorus in, or just outside, each primary costal areole, generally close to costa, at most halfway to margin, elongate, superficial or slightly immersed, solitary on a connective vein or on crossing of a vein and a connective vein, or just outside a connective vein, 38 mm, paraphyses present, clathrate and peltate.
· Rocky places in forest understories; 15002700 m. Guizhou, Sichuan, Yunnan.
三叉扇蕨 san cha shan jue
Polypodium triglossum Baker, Bull. Misc. Inform. Kew 1898: 232. 1898; Selliguea triphylla H. Christ; Neolepisorus triglossus (Baker) Ching.
Rhizome creeping; scales densely set, ferrugineous, broadly lanceolate, margin minutely toothed. Fronds not or slightly dimorphic; stipe brown, slender, 3045 cm, glabrous; lamina trisect into ascending lanceolate segments, middle one up to 40 cm, two lateral segments somewhat shorter, 45.5 cm wide, gradually tapering toward both ends, herbaceous/papery, abaxially pale and with sparse scales near costa abaxially, adaxially green and glabrous; scales adpressed, dark brown, peltate, ovate-cuspidate, membranous; lamina margins entire or slightly undulate; venation prominent, lateral veins parallel, erect-spreading, extending almost to margin, intermediate veinlets anastomosing copiously into small hexagonal areolae with free included veinlets; sori small, oblong, superficial, uniseriate or tending to be biseriate along costa between main veins; paraphyses present, clathrate and peltate.
· Rocky places in forest understories. Yunnan.
Neocheiropteris triglossa is very rare and may be extinct in the field. It is perhaps a natural hybrid between N, palmatopedata and Lepisorus macrosphaerus?
毛鳞蕨属 mao lin jue shu
Rhizome thick, stiff, climbing, densely scaly when young, later glabrescent; scales adpressed, brown, orbicular, peltate, clathrate, with a tuft of stiff long hairs at middle on adaxial side, easily caducous. Fronds clustered to distant, subsessile or with stipe; lamina simple, lanceolate, or loriform, widest at middle, herbaceous or thickly stiffly papery, green or yellow green, glabrous, margin entire or undulate; midrib prominent, veinlets anastomosing, distinct, free included veinlets ending in distinct ovate hydathodes adaxially. Sori round, superficial, or slightly immersed, 13 irregular rows on each side of midrib; paraphyses peltate, clathrate.
One variable species widely distributed from Indonesia and Malesia to E Himalaya, Indochina, and S China.
毛鳞蕨 mao lin jue
Polypodium normale D. Don, Prodr. Fl. Nepal.: 1. 1825; Lepisorus chapaensis C. Christensen & Tardieu; Microsorum normale (D. Don) Ching; Neocheiropteris normalis (D. Don) Tagawa; Neolepisorus normalis (D. Don) Ching; Phymatodes subnormalis Nakai; Pleopeltis normalis (D. Don) T. Moore; P. subnormalis Alderwerelt; Polypodium longifrons Wallich ex Hooker & Greville; P. maculosum H. Christ; Tricholepidium angustifolium Ching; T. angustifolium var. falcato-lineare Ching; T. chapaense (C. Christensen & Tardieu) Ching; T. intermedium Ching; T. lanceolatum Ching & S. K. Wu; T. mutense Ching & S. K. Wu; T. pteropodum Ching; T. subnudum Ching & S. K. Wu; T. tibeticum Ching & S. K. Wu; T. venosum Ching.
Rhizome thick, creeping; scales adpressed, brown or pale brown, orbicular, with a tuft of hairs at middle on adaxial side; fronds close or distant; stipe thick, short or frond subsessile, scaly at base; lamina green or yellow-green, lanceolate to loriform, 3560 Χ 24 cm, widest at middle, herbaceous or thickly papery, glabrous, margin entire or undulate; midribs prominent, veins forming 2 or 3 irregular areoles on each side of midrib, distinct, free included veinlets ending with hydathodes, veinlets at margin of laminae free; sori round, superficial, or slightly immersed, 13 irregular rows on each side of midrib; paraphyses peltate, clathrate.
Climbing on rocks or tree trunks in wet forests; 9002600 m. Guangxi, Xizang, Yunnan [Bhutan, India, Indonesia, Malaysia, Myanmar, Nepal, Thailand, Vietnam].
盾蕨属 dun jue shu
Plants small to medium-sized; rhizome long-creeping; scales pseudopeltate, sometimes peltate, ovate to lanceolate, sometimes orbicular, margin entire or denticulate. Fronds distant, monomorphic; stipe long, scaly; lamina usually simple, entire, sometimes lobed or irregularly lobed, or hastate, herbaceous or papery, scaly; veins anastomosing, areoles regular or irregular, simple to complex, free included veinlets simple or forked; sori in one or two rows on either side of midrib, or in one row between two lateral veins, oblong or rounded to confluent, sometimes linear or slightly irregular; paraphyses peltate, clathrate.
Species number uncertain: northeastern India east to Japan and the Philippines; three species in China.
Terrestrial or on rocks.[??comment]
1a...... Lamina broader at or near base, gradually narrowed to apex ............................................................... 1. N. ovatus
1b..... Fronds broader at or near middle, gradually narrowed to base.
2a...... Lateral veins prominent, smaller veins obscure; spore surface shallowly wavy ........................ 2. N. ensatus
2b..... Veins prominent; spore surface irregularly rugose ............................................................................ 3. N. minor
卵叶盾蕨 luan ye dun jue
Pleopeltis ovata Beddome, Ferns Brit. Ind. t. 157. 1865/6 [not T. Moore, Ind. 78. 1857]; Neolepisorus basicordatus P. S. Wang; N. crenatus S. F. Wu; N. dengii Ching & P. S. Wang; N. emeiensis Ching & K. H. Shing; N. lancifolius Ching & K. H. Shing; N. ovatus (Beddome) Ching & K. H. Shing [??see IPNI]; N. phyllomanes (H. Christ) Ching; N. sinensis Ching; N. tsaii Ching & K. H. Shing; Polypodium phyllomanes H. Christ.
Rhizome creeping; scales ovate-lanceolate, margin sparsely denticulate, long acuminate. Fronds distant; stipe 1020 cm, densely scaly; lamina irregularly pinnatifid, 3-lobed, ovate-lobed, or ovate, 712 cm wide, base rounded, margin entire, apex acuminate; thickly papery, abaxial surface scaly, adaxial surface glabrous; midrib prominent, lateral veins obvious, veinlets anastomosing, free included veinlets forked; sori round, in 24 irregular rows on each side or midrib; covered by paraphyses when young.
Limestone areas, forests; ca. 1500 m. Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan. Yunnan, Zhejiang [Vietnam].
Neolepisorus ovatus is a variable species, with fronds that are sometimes irregularly lobed. Four forms have been proposed. [NO FORMS IN FOC]
[change all of the following into a comment]
1a. Lamina with yellow stripe among lateral veins ........................................................................................ f. truncatus
1b. Lamina without yellow stripe among lateral veins.
2a. Lamina entire, ovate ................................................................................................................................ f. ovatus
2b. Lamina trilobate or irregularly pinnatifid.
3a...... Lamina trilobate or ovate-trilobate, irregularly pinnatifid below middle, lobes lanceolate f. deltoideus
3b...... Lamina trilobate, irregularly deeply pinnatifid, lobes narrowly lanceolate ................... f. doryopteris
盾蕨 (原变型) dun jue (yuan bian xing)
Rhizome creeping; scales ovate-lanceolate, margin sparsely denticulate, long acuminate. Fronds distant; stipe 1020 cm, densely scaly; lamina ovate, 712 cm wide, base rounded, entire, apex acuminate; texture thickly papery, scaly on abaxial surface, glabrous on adaxial surface; midrib prominent, lateral veins obvious, veinlets anastomosing, free included veinlets forked; sori round, 24 irregular rows on each side or midrib; covered by paraphyses when young.
* Anhui, Chongqing, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangsu, Jiangxi, Sichuan, Yunnan, Zhejiang.
The name Polypodium ovatum was first used by Wallich; W. J. Hooker & Greville did validate it (Icon. Filic. 1: t. 41. 1827) but this was a later homonym of P. ovatum Thunberg (1768). T. Moore published Pleopeltis ovata (Wallich) T. Moore but this is a nom. nud. as there is reference only to Wallich, which lacks any description, and no mention of Hooker & Greville; Beddome provided a plate with analysis and reference to Wallich ex Hooker & Greville.
三角叶盾蕨 san jiao ye dun jue
Polypodium deltoideum Baker, J. Bot. 26: 230. 1880 [not Swartz 1788, nor Liebm. 1849]; Neolepisorus dengii Ching & P. S. Wang f. hastatus Ching & P. S. Wang; N. emeiensis Ching & K. H. Shing f. dissectus Ching & K. H. Shing; N. ovatus f. monstrosus Ching & K. H. Shing.
Lamina deltoid with irregular lobes at lower part.
· Forests. Chongqing, Guizhou, Sichuan.
蟹爪盾蕨 xiθ zhua dun jue
Polypodium phyllomanes H. Christ var. doryopteris H. Christ, Bull. Acad. Int. Gιogr. Bot. 11: 214. 1902; Neocheiropteris phyllomanes (H. Christ) Ching var. doryopteris (H. Christ) Ching; Neolepisorus phyllomanes (H. Christ) Ching var. doryopteris (H. Christ) Ching.
Lamina broadly ovate, bipinnatifid at base, lobes linear-lanceolate.
· Guizhou.
截基盾蕨 jie ji dun jue
Neolepisorus truncatus Ching & P. S. Wang, Acta Phytotax. Sin. 21: 270. 1983; N. truncatus f. laciniatus Ching & K. H. Shing.
Lamina ovate-lanceolate, undivided, base truncate, with yellow line between lateral veins.
Limestone areas; ca. 1500 m. Guangxi, Guizhou [Vietnam].
Neolepisorus ovatus f. truncatus is a beautiful plant.
盾蕨 dun jue
Polypodium ensatum Thunberg, Trans. Linn. Soc. London 2: 341. 1794; Microsorum ensatum (Thunberg) H. Itτ; Neocheiropteris ensata (Thunberg) Ching f. izuensis (Sa. Kurata & Satake) Serizawa; N. ensata var. izuensis Sa. Kurata & Satake; N. ensata f. undulatodentata Sugimoto [??not in Tropicos]; Neolepisorus ensatus f. monstrifera Tagawa; N. ensatus f. platyphyllus (Tagawa) Ching & K. H. Shing; N. ensatus var. platyphyllus Tagawa.
Rhizome long-creeping, 35 mm in diam.; scales pale brown, pseudopeltate, ovate to ovate-lanceolate, 26 Χ 11.5 mm, membranous, clathrate, with irregular projections at margin, acuminate; fronds distant; stipe 2030 cm; lamina lanceolate to broadly lanceolate, 2050 Χ 46 cm, broadest at middle, papery, with small adpressed scales on both surfaces when young, cuneate at base, acuminate at apex; veins copiously anastomosing, free included veinlets forked; sori round to oblong, arranged in (1 or)24 rows on each side of midribs, covered with small peltate paraphyses when young.
Guizhou, Sichuan, Taiwan, Yunnan [NE India, Japan, S Korea, Philippines].
小盾蕨 xiao dun jue
Neolepisorus tenuipes Ching & K. H. Shing.
Rhizome long-creeping, ca. 2 mm in diam.; scales brown, broadly lanceolate, acuminate; fronds distant; stipe slender, 1.59 cm, ca. 1 mm in diam., scaly; lamina simple, pale green, ovate to ovate-lanceolate, 618 Χ 23.5 cm, thin, papery, base cuneate, apex acuminate; veins copiously anastomosing [; sori large, round, 1–2 rows between two lateral veins, covered with paraphyses when young].
· Shaded forests, on limestone rocks; 4001600 m. SE Yunnan, ?Zheijiang.
The record of Neolepisorus minor from Zheijiang is based on the treatment of the genus in Fl. Zhejiang (1: 320. 1993).